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Yoshida - 1981 - Fundamentals of Rice Crop Science

Yoshida - 1981 - Fundamentals of Rice Crop Science

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242 FUNDAMENTALS OF RICE CROP SCIENCE<br />

will then examine the so-called high yielding cultivation technology.<br />

7.5.1. Physiological requirements for high yields<br />

Present knowledge <strong>of</strong> crop photosynthesis, mineral nutrition, and yield components<br />

indicates several requirements for high rice yields:<br />

• The variety must have a short, stiff culm.<br />

• The most desirable leaf arrangement is that in which the erect upper leaves<br />

graduate down to droopy leaves at low canopy levels.<br />

• A LAI <strong>of</strong> 5–6 is necessary to achieve maximum crop photosynthesis during<br />

the reproductive stage.<br />

• During ripening, the LAI will decrease as grain filling proceeds and leaf<br />

senescence occurs. While this is a normal event, it is important to maintain as<br />

many active, green leaves as possible until the linear phase <strong>of</strong> grain growth is<br />

completed.<br />

• Planting time should be chosen so that the crop is exposed to high solar<br />

radiation during the reproductive or ripening stages, whichever is more<br />

critical to yield.<br />

• Yield components must meet the requirements for a target yield.<br />

• All the essential nutrients must be supplied to meet the crop’s requirements.<br />

Nitrogen is the most important. The amount <strong>of</strong> nitrogen required to produce 1<br />

t rough rice is about 20 kg and it is independent <strong>of</strong> yield levels. Thus, the<br />

amount <strong>of</strong> nitrogen needed will increase as yield increases. A similar consideration<br />

should apply to other nutrients.<br />

• Nitrogen absorption after heading will assume an increased importance when<br />

high yields are achieved by increasing the harvest index.<br />

During the ripening, about 70% <strong>of</strong> the nitrogen absorbed by the straw will<br />

be translocated to the grain. Nitrogen content <strong>of</strong> the grain tends to be<br />

maintained at a certain percentage. When a larger number <strong>of</strong> grains relative to<br />

the size <strong>of</strong> vegetative parts is produced, more nitrogen will be needed to<br />

support grain growth, and there will be a sharper drain in leaf nitrogen<br />

content. Some grains may suffer from nitrogen shortages.<br />

To produce high yields, it is essential to maintain the level <strong>of</strong> leaf nitrogen<br />

required for high photosynthetic activity. This requirement can be met when<br />

nitrogen absorption by a crop is continued after heading, or when the higher<br />

nitrogen content <strong>of</strong> the vegetative parts is attained before heading so that the<br />

nitrogen absorbed by heading is sufficient for ripening. The Japanese experience<br />

suggests that continuous absorption <strong>of</strong> nitrogen after heading is important<br />

for high yielding crops (Matsushima 1969, Murayama 1967).<br />

7.5.2. Soil culture vs water culture<br />

Water culture has been extensively used in studies <strong>of</strong> the mineral nutrition <strong>of</strong><br />

plants. Compared with soil culture as a rooting medium, water culture provides the<br />

following features:<br />

• Amounts <strong>of</strong> all the essential nutrients and the time <strong>of</strong> their supply can be<br />

controlled.

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