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Yoshida - 1981 - Fundamentals of Rice Crop Science

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238 FUNDAMENTALS OF RICE CROP SCIENCE<br />

anthesis may cause high percentages <strong>of</strong> sterility.<br />

f. Strong winds. Strong winds can cause sterility at flowering by desiccating the<br />

plant, impair grain growth by causing mechanical damage to the grain surface,<br />

and cause the crop to lodge. A hot, dry air called foehn frequently causes white<br />

head, particularly when it occurs during panicle exsertion (Hitaka and Ozawa<br />

1970). Strong winds from the coast <strong>of</strong>ten contain brackish water and cause<br />

sterility.<br />

g. Soil salinity. High salinity in the soil causes a high percentage <strong>of</strong> sterility.<br />

h. Drought. Drought at flowering <strong>of</strong>ten causes high percentages <strong>of</strong> sterility in<br />

rainfed lowland and upland rices.<br />

7.3.4. Grain weight<br />

Grain size is rigidly controlled by hull size. When a piece <strong>of</strong> vinyl film was placed<br />

inside the palea and lemma at anthesis, grain filling proceeded normally but the<br />

grain weight decreased from 22 mg (untreated) to 18 mg (treated) (Matsushima<br />

1970).<br />

Under most conditions, the 1,000-grain weight <strong>of</strong> field crops is a very stable<br />

varietal character (see Table 2.11, Fig. 2.3, and Soga and Nozaki 1957). The CV<br />

<strong>of</strong> 1,000-grain weight ranged from 2.2 to 4.4%, with a mean value <strong>of</strong> 3.3% for<br />

data collected from 9 varieties for 19 years at Konosu, Japan. The mean CVs <strong>of</strong><br />

panicle number and yield were 12.4% and 14.1%, respectively (Matsushima<br />

1970).<br />

Heavy shading before heading changed the hull size and decreased 1,000-grain<br />

weight from about 26 to 21 g (Matsushima 1970). As will be discussed later, CO 2<br />

enrichment also affects the grain weight to some extent.<br />

7.4. Frequency curve <strong>of</strong> individual grain weights <strong>of</strong> Koshiji-wase at<br />

30 days after heading (drawn from data <strong>of</strong> Nakayama 1969).

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