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Narcissus and Daffodil

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Production of <strong>Narcissus</strong> bulbs 109<br />

Production from seed<br />

Production from seed is, of course, the method used by breeders, <strong>and</strong> is also<br />

important for species that form few offsets. <strong>Narcissus</strong> breeders may sow seed soon<br />

after collection, either outdoors or in a cold-frame, <strong>and</strong> germination occurs<br />

somewhat unevenly in spring. Caldwell <strong>and</strong> Wallace (1955) reported that, under<br />

natural conditions, seed of <strong>Narcissus</strong> pseudonarcissus germinated naturally in<br />

November or December, while Wells (1989) stated that, when seed is sown outdoors<br />

in early May soon after collection, N. bulbocodium types start to germinate in<br />

late-August <strong>and</strong> germination continues through the winter, with N. pseudonarcissus<br />

offspring perhaps not germinating until early spring. Under natural conditions,<br />

summer drought may induce dormancy in N. pseudonarcissus (Barkham, 1980).<br />

Rees (1972) stated that narcissus seeds have a cold requirement of many weeks,<br />

<strong>and</strong> Linfield <strong>and</strong> Price (1986) germinated seed of commercial cultivars using a<br />

cold treatment of 12 weeks at 12 °C. Thompson (1977) reported that, in N. bulbocodium,<br />

conditioning imbibed seed for 7 weeks at 26 °C (but not at 6 or 16 °C) gave<br />

rapid germination when subsequently moved to 5–16 °C. This suggested that seed<br />

should be sown in a warm glasshouse <strong>and</strong> moved to cooler conditions after 2<br />

months. Hanks <strong>and</strong> Mathew (1997) reported that synchronous germination could<br />

be obtained in N. cyclamineus, N. bulbocodium var. citrinus <strong>and</strong> N. pseudonarcissus by<br />

keeping imbibed seed at 25–30 °C for 8–12 weeks then transferring to 15 °C.<br />

There is little information on other aspects of the seed physiology of narcissus,<br />

although Caldwell <strong>and</strong> Wallace (1955) reported that seeds of N. pseudonarcissus<br />

were not light sensitive. Hanks <strong>and</strong> Mathew (1997) reported that controlling seedborne<br />

fungi on N. cyclamineus by disinfection, HWT, etc., was difficult. Under<br />

optimum conditions, it takes 3 years for seed-raised plants of species such as<br />

N. bulbocodium <strong>and</strong> N. tri<strong>and</strong>rus to reach flowering size, 4–5 years in other species<br />

<strong>and</strong> 7–8 years in some cultivars (Koopowitz, 1986; Oldfield, 1989). This long<br />

period needed to produce saleable bulbs is a strong disincentive to commercial<br />

production. Germination <strong>and</strong> growing systems for the commercial production<br />

of <strong>Narcissus</strong> species were investigated by Hanks <strong>and</strong> Mathew (1997), who showed<br />

that seedlings could be successful raised in cellular trays for sale as ‘plug plants’<br />

or for growing on, a method useful for species like N. bulbocodium that reach<br />

flowering size quickly.<br />

CONCLUSIONS<br />

This chapter demonstrates that the methods for growing <strong>and</strong> h<strong>and</strong>ling narcissus<br />

bulbs are well documented, but that there remains considerable scope for changes<br />

in husb<strong>and</strong>ry <strong>and</strong> other practices that might make the crop more cost-effectively<br />

produced when growing for processing, or which would enable it to be grown in<br />

an more environmentally friendly way. Some major research needs are evident,<br />

such as the need for cultivars resistant to base rot <strong>and</strong> virus diseases (or their<br />

vectors), for safer <strong>and</strong> (or) more effective chemicals (e.g., for bulb disinfection <strong>and</strong><br />

the control of large narcissus fly), <strong>and</strong> for the development of mechanical or<br />

robotic bulb h<strong>and</strong>ling <strong>and</strong> sorting to reduce labour inputs. The transformation of<br />

narcissus bulb production seen in the UK in recent decades has demonstrated

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