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NIS - libdoc.who.int - World Health Organization

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WHO monographs on medicinal plants commonly used in the Newly Independent States (<strong>NIS</strong>)<br />

Description<br />

Crataegus monogyna: a thorny shrub; leaves bright green with 3 or 5 acute<br />

lobes, deeper and further apart than those of C. laevigata. Flowers,<br />

grouped <strong>int</strong>o branchy corymbs, have 5 triangular sepals, 5 white petals,<br />

and an androecium of 15–20 stamens inserted on the edge of a monocarpellate,<br />

brownish-green receptacle; floral peduncles and sepals pubescent,<br />

stamen with black anthers and 1 style (1, 9).<br />

Crataegus laevigata: a thorny shrub; twigs glabrescent, brown; leaves<br />

bright green, obovate, with 3, 5 or 7 shallow, obtuse lobes. Flowers,<br />

grouped <strong>int</strong>o branchy corymbs, have 5 triangular sepals, 5 white petals,<br />

and an androecium of 15–20 stamens inserted on the edge of a bi- or tricarpellate<br />

receptacle; floral peduncles and sepals glabrous, stamens with<br />

red anthers and 2–3 styles; fruits deep red, globose or ellipsoid (9, 11).<br />

Plant material of <strong>int</strong>erest: dried leaf with flower<br />

General appearance<br />

Crataegus monogyna: leaves bright green with 3 or 5 acute lobes, deeper<br />

and further apart than those of C. laevigata, with secondary venation<br />

curved outwards. Flowers, grouped <strong>int</strong>o branchy corymbs, have 5 triangular<br />

sepals, 5 white petals, and an androecium of 15–20 stamens inserted<br />

on the edge of a monocarpellate, brownish-green receptacle; floral peduncles<br />

and sepals pubescent, anthers black with 1 style; sepals lanceolate,<br />

acuminate, falling over the ovary after flowering (1, 9).<br />

Crataegus laevigata: leaves bright green with 3, 5 or 7 shallow, obtuse,<br />

converging lobes, with secondary venation curved inward. Flowers,<br />

grouped <strong>int</strong>o branchy corymbs, have 5 triangular sepals, 5 white petals,<br />

and an androecium of 15–20 stamens inserted on the edge of a bi- or tricarpellate<br />

receptacle; floral peduncles and sepals glabrous, stamens with<br />

red anthers and 2–3 styles.<br />

Organoleptic properties<br />

Odour: characteristic, fa<strong>int</strong>; taste: slightly bitter-sweet, astringent (12–15).<br />

Microscopic characteristics<br />

Leaf dorsoventral; cells of upper epidermis polygonal, straight-walled<br />

with striated cuticle, those of lower epidermis more sinuous; anomocytic<br />

stomata on lower epidermis only; covering trichomes on both epidermises<br />

but more numerous on the lower, which are long, tapering, unicellular or<br />

very occasionally uniseriate with 2 cells, walls moderately thickened;<br />

cluster crystals or groups of small prismatic crystals of calcium oxalate in<br />

92

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