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NIS - libdoc.who.int - World Health Organization

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WHO monographs on medicinal plants commonly used in the Newly Independent States (<strong>NIS</strong>)<br />

Geographical distribution<br />

Indigenous to Central America, it is now cultivated worldwide as green<br />

fodder or as a cereal crop (2, 5, 20–23).<br />

Description<br />

A robust monoecious annual plant, up to 3 m in height. Roots, fasciculate;<br />

often with prop roots from the lower nodes. Stems, erect, culm-<strong>int</strong>ernodes<br />

solid, smooth. Leaves, alternate, with sheathing base, ciliate ligule about 5 mm<br />

long, broadly linear or lanceolate blade, upper surface hairy, lower surface<br />

hairless, acuminate, parallel-veined, 25–100 cm long, 2–10 cm wide. Flowers,<br />

unisexual. Inflorescences, terminal panicle of male flowers, and spadix of female<br />

flowers. Male inflorescence (tassel) is a terminal large spreading panicle,<br />

about 30 cm long, formed by spike-like racemes of staminate spikelets. The<br />

spikelets are in pairs, one sessile, the other pedicelled, 8–12 mm long, awnless,<br />

2-flowered, florets both male; glumes papery, equal, enclosing florets. Female<br />

inflorescence (spadeces or ears) variable in size and shape, borne on short<br />

branch with several short <strong>int</strong>ernodes with a papery sheath at each node, these<br />

form the husk and enclose the thick central axis (cob) on which the spikelets<br />

are arranged in 8–16(30) longitudinal rows; spikelets in pairs, 2-flowered,<br />

both sessile, awnless, the lower floret small, rarely female, the upper one female;<br />

glumes broad, rounded or notched at apex, fleshy towards base; styles<br />

projecting from apex of ear and look like tufts of hair, at first green, later red<br />

or yellow. Fruit, caryopsis with adherent pericarp; exposed between gaping<br />

lemma and palea at maturity; variable as to size, shape and colour (usually<br />

yellow but can be whitish or darker, almost black) (2, 5, 24–27).<br />

Plant material of <strong>int</strong>erest: dried styles and stigmas<br />

General appearance<br />

Slender, yellowish or brownish, filamentous styles, 5–20 cm long, exhibiting<br />

slender bifid stigmas 0.5–3.0 mm long. Viewed with a hand lens, they<br />

appear flattened and ribbon-like, or grooved and curled up, with spreading<br />

trichomes (1, 28, 29). In the cut form, the drug is made up of 5–10-mm<br />

long filamentous, channelled pieces of stigmata, pale yellowish or brownish<br />

red in colour (5). According to the USSR pharmacopoeia, the cut drug<br />

is composed of fragments of styles and stigmas having a length less than<br />

7 mm, they are brownish, brownish red or light yellow in colour (1).<br />

Organoleptic properties<br />

Odour: slight, fa<strong>int</strong>, characteristic; taste: insipid, slightly mucilaginous,<br />

somewhat sweetish (1, 5, 28, 29).<br />

424

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