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Grasslands of the World.pdf - Disasters and Conflicts - UNEP

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<strong>Grassl<strong>and</strong>s</strong> <strong>of</strong> South Africa 97<br />

Nama-karoo<br />

The Nama-karoo biome covers much <strong>of</strong> <strong>the</strong> central <strong>and</strong> western regions <strong>of</strong> <strong>the</strong><br />

country. The biome is dominated by a steppe-type vegetation , comprising a<br />

mixture <strong>of</strong> shrubs, dwarf shrubs <strong>and</strong> annual <strong>and</strong> perennial grasses. The biome<br />

is associated with <strong>the</strong> moderate rainfall regions (250-450 mm per annum) <strong>and</strong><br />

is suited to commercial sheep <strong>and</strong> goat production. The summer seasonality <strong>of</strong><br />

<strong>the</strong> rainfall in <strong>the</strong> eastern parts <strong>of</strong> <strong>the</strong> biome means that <strong>the</strong>re is <strong>of</strong>ten abundant<br />

grass production during <strong>the</strong> growing season. Graziers attempt to optimize<br />

production by sparing or resting grassy dwarf shrubl<strong>and</strong> in <strong>the</strong> wet season.<br />

Herbivory by domestic livestock during <strong>the</strong> growing season has been shown<br />

to reduce grass cover <strong>and</strong> promote <strong>the</strong> growth <strong>of</strong> larger shrubs (species <strong>of</strong><br />

Rhus , Acacia <strong>and</strong> Euclea ) <strong>and</strong> dwarf shrubs. In <strong>the</strong> winter months, <strong>the</strong> dwarf<br />

shrubs maintain <strong>the</strong>ir crude protein at around 8 percent, providing excellent<br />

forage. The nutrient-rich substrata provided by <strong>the</strong> mudstones, s<strong>and</strong>stones <strong>and</strong><br />

dolerites mean that this production can be considered sustainable . There were<br />

earlier suggestions that large-scale structural transformations were taking place<br />

in this biome (Acocks, 1964), with <strong>the</strong> dwarf shrubs supposedly spreading into<br />

<strong>the</strong> adjoining grassl<strong>and</strong>s <strong>of</strong> central Free State. This process has not continued in<br />

<strong>the</strong> manner envisaged <strong>and</strong> <strong>the</strong> relatively high rainfall <strong>of</strong> <strong>the</strong> 1990s has promoted<br />

grass production in <strong>the</strong> eastern portions <strong>of</strong> <strong>the</strong> biome. In <strong>the</strong> western portions<br />

<strong>of</strong> <strong>the</strong> biome, <strong>the</strong>re is alarming evidence <strong>of</strong> woody encroachment , with two<br />

species in particular (Acacia mellifera <strong>and</strong> Rhigozum trichotomum ) increasing<br />

in density <strong>and</strong> cover in regions with a long history <strong>of</strong> domestic herbivory.<br />

Production <strong>of</strong> mutton <strong>and</strong> fibre continues to thrive in <strong>the</strong> Nama-karoo. During<br />

<strong>the</strong> recent past <strong>the</strong>re has also been an increase in <strong>the</strong> area <strong>of</strong> l<strong>and</strong> set aside for<br />

informal conservation , with many farmers capitalizing on <strong>the</strong> unique l<strong>and</strong>scapes<br />

<strong>and</strong> indigenous fauna <strong>of</strong> <strong>the</strong> biome to develop ecotourism operations.<br />

Important indigenous herbivores, which contribute to red meat production,<br />

include springbok, blesbok, kudu, gemsbok <strong>and</strong> wildebeest.<br />

Thicket<br />

The thicket biome occurs in <strong>the</strong> drainage lines <strong>and</strong> ridges <strong>of</strong> <strong>the</strong> sou<strong>the</strong>astern<br />

coastal region <strong>and</strong> inl<strong>and</strong> to <strong>the</strong> Great Escarpment. The thicket comprises a<br />

dense cover <strong>of</strong> succulent shrubs, woody shrubs <strong>and</strong> small trees, with a height<br />

<strong>of</strong> 1.5–3.0 m. The woody shrubs are multistemmed, seasonally-deciduous, C3<br />

plants. In <strong>the</strong> xeric portions <strong>of</strong> <strong>the</strong> thicket biome (300–450 mm/yr precipitation ),<br />

<strong>the</strong>re is a large component <strong>of</strong> crassulacean acid metabolism (CAM )-type leaf<br />

succulents (e.g. Portulacaria afra ), stem succulents (Euphorbia spp.) <strong>and</strong> many<br />

species <strong>of</strong> small succulent shrubs (e.g. species <strong>of</strong> Aloe <strong>and</strong> Crassula ). Important<br />

grass species that contribute significantly to cattle production include Panicum<br />

maximum , P. deustum , Digitaria eriantha <strong>and</strong> Setaria sphacelata . Although<br />

<strong>the</strong> thicket biome does not contain extensive grassl<strong>and</strong>s, clearing <strong>of</strong> thicket is<br />

carried out by both freehold <strong>and</strong> communal graziers to promote grassl<strong>and</strong>s.

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