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Grasslands of the World.pdf - Disasters and Conflicts - UNEP

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134<br />

<strong>Grassl<strong>and</strong>s</strong> <strong>of</strong> <strong>the</strong> world<br />

Semi-deserts <strong>and</strong> shrub steppes<br />

Semi-deserts <strong>and</strong> shrub steppes (Units Kg 11 <strong>and</strong> Kf 11 in Figure 4.5) exhibit<br />

similar latitudinal extents; toge<strong>the</strong>r, both variants <strong>of</strong> semi -desert cover 22 percent<br />

<strong>of</strong> <strong>the</strong> region, with plant communities <strong>of</strong> low diversity, including, on average,<br />

19 plant species (Golluscio, León <strong>and</strong> Perelman, 1982). The MAP <strong>of</strong> much <strong>of</strong><br />

<strong>the</strong> area is less than 150 mm. Patagonian semi-deserts are less productive than<br />

shrub steppes; ANPP calculated from NDVI -I values by Paruelo, Jobbágy <strong>and</strong><br />

Sala (1998b) are 450 <strong>and</strong> 390 kg/ha/yr for units Kg 11 <strong>and</strong> Kf 11, respectively<br />

(Table 4.1). From a species composition st<strong>and</strong>point, Kf 11 is <strong>the</strong> most typical<br />

<strong>and</strong> less degraded semi-desert type , in spite <strong>of</strong> being less productive (Soriano,<br />

1983). Dwarf shrubs with cushion habits are typical <strong>of</strong> this vegetation type in <strong>the</strong><br />

Kg 11 <strong>and</strong> Kf 11 units. Nassauvia glomerulosa , N. ulicina <strong>and</strong> Chuquiraga aurea<br />

are dominant, accompanied by o<strong>the</strong>rs such as Chuquiraga kingii , Brachyclados<br />

caespitosus <strong>and</strong> Perezia lanigera (León et al., 1998). Grasses such as Stipa<br />

humilis , S. ibarii , S. ameghinoi <strong>and</strong> shrubs such as Chuquiraga avellanedae ,<br />

Schinus polygamus <strong>and</strong> Lycium chilense are secondary species in semi-desert<br />

plant communities . Some co-dominant species occur, but only in <strong>the</strong> Kg 11<br />

semi-deserts: dwarf shrubs such as Azorella caespitosa , Mullinum microphyllum<br />

<strong>and</strong> Frankenia sp. , grasses such as Poa dusenii , P. ligularis <strong>and</strong>, less frequently,<br />

Stipa neaei , <strong>and</strong> shrubs such as Junellia tridens , which occur in clumps on paleodepressions<br />

<strong>and</strong> natural drainage networks (León et al., 1998; Roig, 1998).<br />

Shrub-grass <strong>and</strong> grass-shrub steppes<br />

Shrub-grass <strong>and</strong> grass-shrub steppes span almost <strong>the</strong> entire latitudinal extent<br />

<strong>of</strong> Patagonia <strong>and</strong> cover approximately 20 percent <strong>of</strong> <strong>the</strong> arid area (Units Jf 11<br />

<strong>and</strong> Jd 12 in Figure 4.5). According to Paruelo, Jobbágy <strong>and</strong> Sala (1998b), both<br />

types have similar ANPP levels (650 kg/ha/yr), although NDVI values peak<br />

somewhat later in <strong>the</strong> growing season in grass-shrub steppes . Vegetation cover<br />

<strong>of</strong> <strong>the</strong> shrub-grass steppes is 47 percent, with plant communities that contain<br />

about 26 vascular plant species (Golluscio, León <strong>and</strong> Perelman, 1982). Adesmia<br />

campestris , Mullinum spinosum , Senecio filaginoides , Berberis heterophylla ,<br />

Colleguaya integerrima , Trevoa patagonica <strong>and</strong> Schinus polygamus are <strong>the</strong><br />

most conspicuous shrubs (León et al., 1998). The most important grasses are<br />

Stipa speciosa , S. humilis , Poa ligularis , P. lanuginosa , Festuca argentina <strong>and</strong><br />

F. pallescens , <strong>and</strong> occur with sedges <strong>of</strong> <strong>the</strong> genus Carex (León et al., 1998).<br />

Grass -shrub steppes are a transition between <strong>the</strong> shrub steppes <strong>and</strong> grass<br />

steppes on <strong>the</strong> one h<strong>and</strong> <strong>and</strong> <strong>the</strong> grass-steppes <strong>and</strong> <strong>the</strong> Nothophagus subantarctic<br />

forest ecotone on <strong>the</strong> o<strong>the</strong>r (Figure 4.5) (Roig, 1998). Paruelo, Jobbágy <strong>and</strong><br />

Sala (1998b) combine <strong>the</strong> latter (Chiliotrichum diffusum grass-shrub steppes)<br />

<strong>and</strong> <strong>the</strong> steppe-forest ecotones in sou<strong>the</strong>rn Santa Cruz <strong>and</strong> nor<strong>the</strong>rn Tierra<br />

del Fuego into a single biozone (unit Ha 12). Physiognomically, <strong>the</strong>se are<br />

different domains; <strong>the</strong>y may exhibit similar functional attributes <strong>and</strong> should<br />

be significantly more productive than <strong>the</strong> grass-shrub steppes at <strong>the</strong> opposite

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