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Grasslands of the World.pdf - Disasters and Conflicts - UNEP

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The Russian Steppe 391<br />

pratensis dominant. 6. Association group: various admixtures <strong>of</strong> Alopecurus<br />

pratensis. 7. Associations.<br />

Sukachev’s classification is “phytocoenological”, whereas Ramenskii’s is<br />

“phytotopological”, <strong>and</strong> <strong>the</strong> latter’s example was followed by Dmitriev (1948)<br />

<strong>and</strong> Chugunov (1951). However, <strong>the</strong> distinction is <strong>of</strong>ten blurred. Ramenskii’s<br />

seral stages <strong>of</strong> plant succession, although not coined as such by him or his successors,<br />

are classical examples <strong>of</strong> linear Clementsian succession.<br />

Conversely, bare ground is colonized by ruderals, which give place to<br />

seral grassl<strong>and</strong> stages as organic matter accumulates <strong>and</strong> <strong>the</strong>se are eventually<br />

replaced by taller bunch grasses. Clements (1916) ideas proved applicable not<br />

only in USA , but also in Canada (Coupl<strong>and</strong>, 1979, whose work used to be<br />

quoted in Russia ) <strong>and</strong> in East <strong>and</strong> South Africa (Phillips, 1929). It is doubtful<br />

if it ever was Clements’ aim to apply <strong>the</strong> succession model to all situations<br />

liable to transition, or to claim that climax vegetation was <strong>the</strong> most desirable<br />

or most productive from <strong>the</strong> agricultural point <strong>of</strong> view in all situations. Plant<br />

succession can be studied with <strong>the</strong> aim <strong>of</strong> identifying <strong>the</strong> preferred seral stages<br />

with desirable composition. As we will see in Russia, many instances can also<br />

be found in which some <strong>of</strong> <strong>the</strong> seral stages (fallow l<strong>and</strong>) are considerably more<br />

diverse <strong>and</strong> productive in herbage than <strong>the</strong> climax vegetation itself. It goes too<br />

far, however, to regard this as counter-evidence for <strong>the</strong> succession model. As in<br />

<strong>the</strong> Great Plains <strong>of</strong> USA, <strong>the</strong> climax grasses <strong>of</strong> <strong>the</strong> Russian steppe are productive<br />

<strong>and</strong> acceptable to livestock; <strong>the</strong>y provide ground cover to protect soil <strong>and</strong><br />

are effective plants in utilizing environmental growth factors to fix carbon <strong>and</strong><br />

to cycle nutrients.<br />

The interplay <strong>of</strong> biotic, climatic <strong>and</strong> edaphic components <strong>of</strong> <strong>the</strong> environment<br />

is relevant, as <strong>the</strong>se modify <strong>the</strong> dynamics <strong>of</strong> grassl<strong>and</strong> communities . Efforts at<br />

grassl<strong>and</strong> improvement are directed at manipulating <strong>the</strong> botanical composition<br />

to encourage <strong>the</strong> more desirable species <strong>and</strong> suppress those less desirable. A basic<br />

thrust in current grassl<strong>and</strong> improvement has been <strong>the</strong> comparison <strong>of</strong> <strong>the</strong> current<br />

site condition with what it ought to be, i.e. site potential . Acknowledgement <strong>of</strong><br />

various stable vegetation states at a particular site widens <strong>the</strong> scope for opportunistic<br />

management that is responsive to abiotic events <strong>and</strong> that is not bound<br />

under all circumstances by doctrines that abhor fire or that only value stocking<br />

rates that are moderate.<br />

BOTANICAL CONDITION (ECOLOGICAL MONITORING )<br />

Few are <strong>the</strong> instances where ecological techniques can have such an impact<br />

on practical management decisions in agriculture as those developed in<br />

grassl<strong>and</strong> science. Botanical assessment has proved a more reliable <strong>and</strong> efficient<br />

criterion <strong>of</strong> condition , <strong>and</strong>, <strong>the</strong>refore, <strong>of</strong> productivity <strong>of</strong> grassl<strong>and</strong>, than yield<br />

measurements <strong>the</strong>mselves. Given <strong>the</strong> enormous tasks <strong>of</strong> natural resource<br />

management ahead, it is appropriate here to draw attention to new techniques<br />

<strong>of</strong> botanical grassl<strong>and</strong> surveys.

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