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Grasslands of the World.pdf - Disasters and Conflicts - UNEP

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The changing face <strong>of</strong> pastoral systems in grass-dominated ecosystems <strong>of</strong> eastern Africa 37<br />

persistent <strong>and</strong> withst<strong>and</strong>s severe defoliation <strong>and</strong> grazing . It is <strong>the</strong> dominant<br />

species in natural pastures in many parts <strong>of</strong> <strong>the</strong> eastern Africa n highl<strong>and</strong>s. It<br />

is an invasive secondary species, which can quickly colonize disturbed soil in<br />

cropping areas <strong>and</strong> fallow l<strong>and</strong>, spreading by seeds or stolons, <strong>and</strong> may become<br />

a serious weed in cropl<strong>and</strong> (Boonman, 1993). It shows wide variability, with<br />

three distinct ecotypes classified on leaf width <strong>and</strong> length, stolon size <strong>and</strong> floral<br />

structure (Skerman <strong>and</strong> Riveros, 1990). Several ecotypes have been selected<br />

as commercial cultivars, which have been widely introduced into tropical<br />

highl<strong>and</strong> <strong>and</strong> subtropical areas. It is now widely grown outside its native<br />

distribution <strong>and</strong> is commonly cultivated in <strong>the</strong> Americas. Studies in <strong>the</strong> USA<br />

using starch gel electrophoresis to describe <strong>the</strong> distribution <strong>of</strong> genetic variation<br />

within <strong>and</strong> among introduced populations found a relatively high proportion<br />

<strong>of</strong> polymorphic loci across populations, indicating fixed heterozygosity due to<br />

polyploidy (Wilen et al., 1995). The highl<strong>and</strong> grazing areas <strong>of</strong> P. cl<strong>and</strong>estinum<br />

are <strong>of</strong>ten mixed with P. sphacelatum <strong>and</strong> Eleusine floccifolia . These two grasses<br />

are frequent in overgrazed pastures in <strong>the</strong> highl<strong>and</strong>s <strong>and</strong> mid-altitudes in <strong>the</strong><br />

Rift Valley, but are not palatable (Sisay <strong>and</strong> Baars, 2002) <strong>and</strong> are important for<br />

traditional basket making. Cattle avoid <strong>the</strong>se grasses, which have <strong>the</strong> potential<br />

to become major weeds on upl<strong>and</strong> pastures unless collected for basket making.<br />

Basket making is an important activity <strong>and</strong> source <strong>of</strong> income for rural women<br />

<strong>and</strong> collection <strong>of</strong> <strong>the</strong>se weedy grasses also maintains <strong>the</strong> quality <strong>of</strong> <strong>the</strong> communal<br />

grazing areas <strong>and</strong> grassl<strong>and</strong>s in <strong>the</strong> highl<strong>and</strong>s.<br />

Pennisetum purpureum is a tall, erect, vigorous perennial species that grows<br />

in damp grassl<strong>and</strong>s <strong>and</strong> forest areas up to 2 400 m in Kenya, Tanzania , Ug<strong>and</strong>a<br />

<strong>and</strong> Sudan. Herlocker (1999) recognized this as a vegetation region in Kenya<br />

<strong>and</strong> Ug<strong>and</strong>a, around <strong>the</strong> shores <strong>of</strong> Lake Victoria. Pennisetum purpureum<br />

is widely distributed through sub-Saharan Africa <strong>and</strong> is commonly called<br />

elephant grass or Napier grass, named after Colonel Napier <strong>of</strong> Bulawayo in<br />

Zimbabwe, who promoted its use at <strong>the</strong> start <strong>of</strong> <strong>the</strong> century. It is now widely<br />

used for cut-<strong>and</strong>-carry (where grass is collected by h<strong>and</strong> <strong>and</strong> carried to stallfed<br />

cattle ) for <strong>the</strong> smallholder dairy industry in eastern Africa <strong>and</strong> frequently<br />

produces up to 10–12 t/ha dry matter in rainfed conditions (Boonman, 1993).<br />

Elephant grass is palatable when young <strong>and</strong> leafy. It is fast growing <strong>and</strong> should<br />

be cut <strong>of</strong>ten to avoid its becoming tough <strong>and</strong> unpalatable with a high proportion<br />

<strong>of</strong> stem. Due to its importance in <strong>the</strong> region, considerable research has been<br />

done on elephant grass, including studies on its diversity. Tcacenco <strong>and</strong> Lance<br />

(1992) studied 89 morphological characters on 9 genotypes <strong>of</strong> elephant grass to<br />

determine which characters were most useful for description <strong>of</strong> <strong>the</strong> variation<br />

in <strong>the</strong> species, <strong>and</strong> concluded that variation existed from plant to plant, even<br />

within <strong>the</strong> same accession. A larger collection <strong>of</strong> 53 accessions was characterized<br />

for 20 morphological <strong>and</strong> 8 agronomic characters (Van de Wouw, Hanson<br />

<strong>and</strong> Leuthi, 1999). Again <strong>the</strong> germplasm was found to be very variable, but<br />

accessions could be clustered into six groups with similar morphology. More

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