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Grasslands of the World.pdf - Disasters and Conflicts - UNEP

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98<br />

<strong>Grassl<strong>and</strong>s</strong> <strong>of</strong> <strong>the</strong> world<br />

These grassl<strong>and</strong>s provide high quality, year-round grazing , as <strong>the</strong> shale-derived<br />

soils associated with thicket are rich in nutrients.<br />

Two important thicket types are recognized: succulent thicket <strong>and</strong> mesic<br />

thicket. Succulent thicket occurs under semi -arid conditions (300–450 mm/<br />

yr) <strong>and</strong> is dominated by leaf- <strong>and</strong> stem-succulent shrubs (Portulacaria afra ,<br />

Euphorbia bothae , E. ledienii , E. coerulescens , E. tetragona , E. triangularis ,<br />

numerous Crassula spp. <strong>and</strong> Aloe spp. ). Mesic thicket contains fewer succulents<br />

<strong>and</strong> occurs under higher rainfall (>450 mm/yr) conditions. The cover by woody<br />

shrubs is usually continuous , but <strong>the</strong>y seldom exceed 4 m in height.<br />

Short succulent thicket occurs in <strong>the</strong> low elevation (300–350 m) inl<strong>and</strong> river<br />

valleys <strong>of</strong> <strong>the</strong> Great Fish, Bushmans, Sundays <strong>and</strong> Gamtoos rivers. Here <strong>the</strong><br />

Ecca series mudstones provide a nutrient-rich substratum. The soils are shallow,<br />

comprising arid lithosols derived from <strong>the</strong> basement rock. The valley climate is<br />

hot <strong>and</strong> dry , with extremely high summer maximum temperatures (45°C) <strong>and</strong><br />

low winter minimum temperatures. In <strong>the</strong> Great Fish river valley, <strong>the</strong> structure<br />

<strong>and</strong> composition <strong>of</strong> <strong>the</strong> “pristine” form <strong>of</strong> short succulent thicket is dominated<br />

by Portulacaria afra , Rhigozum obovatum <strong>and</strong> Euphorbia bothae , in small (3–4 m<br />

diameter) clumps. The vegetation in <strong>the</strong> clumps may contain emergant shrubs (e.g.<br />

Boscia oleoides ), but <strong>the</strong>y are usually

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