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keys to the vascular plants of east texas - Botanical Research ...

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7. Leaves straight or nearly so, lacking strongly inrolled margins, usually pale bluish <strong>to</strong> sage<br />

green, conspicuously glaucous, � smooth on both surfaces; leaf margins yellowish, flat<br />

YUCCA/AGAVACEAE<br />

415<br />

__________________________________________________________________________ Y. pallida<br />

5. Leaf margins whitish, usually shredding in<strong>to</strong> prominent white fibers (<strong>the</strong>se <strong>of</strong>ten disappear-<br />

ing late in year); pistils usually 2–3.2 cm long.<br />

8. Inflorescence usually unbranched and raceme-like or with 1 or 2 short, spreading branches<br />

near base (<strong>the</strong>se <strong>of</strong>ten soon deciduous), borne below <strong>to</strong> just above leaf tips (separated<br />

from <strong>the</strong>m by less than its own length <strong>of</strong> naked scape) _____________________________ Y. arkansana<br />

8. Inflorescence a much branched panicle, borne well above tips <strong>of</strong> leaves (separated from<br />

<strong>the</strong>m by nearly its own length or more <strong>of</strong> naked scape).<br />

9. Leaves very slender, 8–15 mm wide, 100–200 per strikingly globose rosette; fruit usually<br />

constricted near middle; <strong>plants</strong> <strong>of</strong> limes<strong>to</strong>ne substrates ___________________________ Y. constricta<br />

9. Leaves usually (6–)15–40 mm wide, ca. 50–85 per rosette (rosette not globose in appearance);<br />

fruit constricted or not so; <strong>plants</strong> <strong>of</strong> sandy substrates.<br />

10. Inflorescences pubescent (rarely glabrous); fruits usually constricted near <strong>the</strong> middle;<br />

<strong>plants</strong> <strong>of</strong> Pineywoods and Post Oak Savannah ____________________________ Y. louisianensis<br />

10. Inflorescences glabrous; fruits usually not conspicuously constricted; <strong>plants</strong> <strong>of</strong> w edge<br />

<strong>of</strong> Blackland Prairie (Dallas Co.) and westward _________________________________ Y. necopina<br />

Yucca aloifolia L., (with leaves like Aloe, Asphodelaceae), DAGGER PLANT, ALOE YUCCA, SPANISH-<br />

BAYONET. Plant large, <strong>to</strong> 3(–7) m tall, very variable morphologically; stems 1–3, unbranched or<br />

sparingly branched, <strong>of</strong>ten growing parallel <strong>to</strong> <strong>the</strong> ground for part <strong>of</strong> <strong>the</strong>ir length; leaves 12–40(–50)<br />

cm long, 2.5–6 cm wide, dark green, rigid, marginally denticulate, rarely with straight fibers;<br />

inflorescences paniculate, glabrous or slightly pubescent; perianth globose, creamy white (can<br />

be tinged with green or purple basally); fruits 3.5–5 cm long, indehiscent, pulpy. Widely cultivated<br />

in East TX, escaping or persisting for decades; old homesteads, dump sites, along railroads,<br />

usually sandy soils but sometimes black clayey soils; Jasper, Liberty, New<strong>to</strong>n, Tyler, and<br />

Walker (Eric Keith, pers. comm.) cos.; native <strong>to</strong> <strong>the</strong> se U.S. from VA s <strong>to</strong> FL w <strong>to</strong> coastal TX (but<br />

only naturalized in East TX). Jun–Aug(–fall). [Y. serrulata Haw.] “Yucca aloifolia has been<br />

widely cultivated, and horticultural forms (or varieties, depending upon <strong>the</strong> source) differ in<br />

<strong>the</strong> striping <strong>of</strong> yellow and white on <strong>the</strong> leaves” (Hess & Robbins 2002). This species is sometimes<br />

mistaken for Y. gloriosa var. recurvifolia, ano<strong>the</strong>r species <strong>of</strong> <strong>the</strong> se U.S. that is also cultivated,<br />

persists, and escapes in <strong>east</strong> Texas; however, that species has longer (40–100 cm long)<br />

mostly recurved, flexible, usually entire leaves and � lea<strong>the</strong>ry fruits.<br />

Yucca arkansana Trel., (<strong>of</strong> Arkansas), ARKANSAS YUCCA, SOAPWEED. Leaves 20–60(–70) cm long,<br />

10–25 mm wide, <strong>the</strong> margins at first white, papery with curly fibers; inflorescence glabrous;<br />

perianth 32–65 mm long, greenish white, globose; capsules ca. 4–7 cm long. Rocky limes<strong>to</strong>ne or<br />

sandy soils, prairies and hillsides; Pineywoods and Gulf Prairies and Marshes w <strong>to</strong> Rolling<br />

Plains and e Edwards Plateau; AR, KS, MO, OK, and TX. Mar–mid-May (this is <strong>the</strong> earliest<br />

bloomer <strong>of</strong> <strong>the</strong> dry-fruited Yuccas in Texas—K. Clary, pers. comm.). [Y. angustissima Engelm. ex<br />

Trel. var. mollis Engelm.] The roots, containing saponins, were used as a soap by both Native<br />

Americans and pioneers and <strong>the</strong> seeds were eaten raw, roasted, or ground in<strong>to</strong> a flour<br />

(Yatskievych 1999). DNA evidence (Clary 1997) suggests that this species is related <strong>to</strong> Y.<br />

louisianensis. m/307<br />

Hybrids <strong>of</strong> Y. arkansana and Y. pallida have been found on limes<strong>to</strong>ne in Dallas (McKelvey<br />

1947), <strong>to</strong> <strong>the</strong> w <strong>of</strong> East TX at Glen Rose in Somervell Co. (Shinners 1958), and recently in Tarrant<br />

County at Tandy Hills Park (BRIT). This latter population <strong>of</strong> hundreds <strong>of</strong> individuals over<br />

a number <strong>of</strong> acres is quite variable, with individuals ranging from much like typical Y.<br />

arkansana <strong>to</strong> those much like Y. pallida as well as a full spectrum <strong>of</strong> intermediates. The <strong>plants</strong><br />

vary from having leaves with curly fibers on <strong>the</strong> margins <strong>to</strong> not so, from having leaf margins

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