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keys to the vascular plants of east texas - Botanical Research ...

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MICROSTEGIUM/POACEAE<br />

977<br />

AA variable C3 genus <strong>of</strong> 60–84 species (depending on inclusion <strong>of</strong> segregates) <strong>of</strong> temperate<br />

regions excluding Australia (Mejía-Saulés & Bisby 2000; Barkworth ined.). The genus is unusual<br />

in having some species (subgenus Melica) with disarticulation below <strong>the</strong> glumes and<br />

o<strong>the</strong>rs (subgenus Bromelica) with disarticulation above <strong>the</strong> glumes (Clay<strong>to</strong>n & Renvoize 1986;<br />

Yatskievych 1999). More specifically, “The North American species fall in<strong>to</strong> two groups. In one,<br />

<strong>the</strong> pedicels are sharply bent and <strong>the</strong> spikelets disarticulate below <strong>the</strong> glumes; in <strong>the</strong> o<strong>the</strong>r, <strong>the</strong><br />

pedicels are straight and disarticulation is above <strong>the</strong> glumes” (Barkworth ined.). (Greek: meli,<br />

honey, classical name for some plant, possibly a species <strong>of</strong> sorghum with sweet sap (Barkworth<br />

ined.), taken up by Linnaeus for this genus) (subfamily Pooideae, tribe Meliceae)<br />

REFERENCES: Boyle 1945; Tucker 1996; Mejía-Saulés & Bisby 2000; Barkworth ined.<br />

1. Panicles usually simple (rarely compound), <strong>the</strong> branches <strong>the</strong>mselves not branched; glumes ca.<br />

equal in length or <strong>the</strong> upper slightly longer; spikelets � flat-<strong>to</strong>pped and triangular in shape;<br />

ligule 1.5 mm or less long; rudiments broadly obovate <strong>to</strong> obconic, truncate (appearing abruptly<br />

cut <strong>of</strong>f ), oriented at a sharp angle <strong>to</strong> <strong>the</strong> rachilla _________________________________________ M. mutica<br />

1. Panicles usually compound, <strong>the</strong> branches <strong>the</strong>mselves branched; upper glume notably longer<br />

than <strong>the</strong> lower; spikelets nei<strong>the</strong>r flat-<strong>to</strong>pped nor triangular in shape; ligule (1–)3–6.5 mm long;<br />

rudiments narrowly obovate or oblong, not truncate, oriented in a � straight line with <strong>the</strong> rachilla<br />

_________________________________________________________________________________ M. nitens<br />

Melica mutica Walter, (blunt, pointless), TWO-FLOWER MELIC, NARROW MELIC. Culms usually<br />

40–80(–100) cm tall from creeping rhizomes; ligule 1.5 mm or less long; leaf blades ca. 2–6 mm<br />

wide; inflorescence 4–16(–25) cm long; spikelets 7–11 mm long; fertile florets usually 2, <strong>the</strong>ir<br />

tips about at <strong>the</strong> same level in <strong>the</strong> spikelet; rudiments broadly obovate <strong>to</strong> obconic, almost clublike.<br />

Forest openings on sandy soils; widespread in Pineywoods, less so in Post Oak Savannah;<br />

also n Gulf Marshes and Prairies and sparsely scattered <strong>to</strong> <strong>the</strong> w; e U.S. from NJ s <strong>to</strong> FL w <strong>to</strong> IA<br />

and TX. Apr–Jun.<br />

Melica nitens (Scribn.) Nutt. ex Piper, (shining), THREE-FLOWER MELIC, TALL MELIC, LADD’S-FAVOR-<br />

ITE. Plant rhizoma<strong>to</strong>us; culms 50–120(–150) cm tall; ligule (1–)3–6.5 mm long; leaf blades 3–11<br />

mm wide; inflorescence usually 10–26 cm long; spikelets 8–12(–15) mm long, much longer than<br />

broad; fertile florets usually 2–3, <strong>the</strong>ir tips at different levels in <strong>the</strong> spikelet. Woodlands, rocky<br />

grasslands; Bexar, Grayson, Limes<strong>to</strong>ne, Travis (BRIT), Austin, Burleson, Comal, and Williamson<br />

(Turner et al. 2003) cos. in w and s parts <strong>of</strong> East TX; mainly w 2/3 <strong>of</strong> TX; e U.S. from PA s <strong>to</strong> GA<br />

w <strong>to</strong> MN and TX, also AZ and NM. Apr–Jun.<br />

MICROSTEGIUM Nees BROWNTOP, SASA GRASS, JAPANESE GRASS<br />

AA genus <strong>of</strong> ca. 15 species (Thieret 2003b) <strong>of</strong> tropical and warm areas <strong>of</strong> <strong>the</strong> Old World, particularly<br />

se Asia. Like all members <strong>of</strong> <strong>the</strong> Andropogoneae, Microstegium is characterized by C4<br />

pho<strong>to</strong>syn<strong>the</strong>sis (Kellogg 2000a). Preliminary molecular evidence suggests that Microstegium<br />

may be polyphyletic and that it is closely related <strong>to</strong> such genera as Miscanthus and Saccharum<br />

(Kellogg 2000a; Ma<strong>the</strong>ws et al. 2002). A relationship <strong>to</strong> Eulalia has also been suggested (Clay<strong>to</strong>n<br />

& Renvoize 1986). (Greek: mikros, small, little, and stego, shelter, cover, possibly in reference<br />

<strong>to</strong> <strong>the</strong> small lemma or glumes) (subfamily Panicoideae, tribe Andropogoneae)<br />

REFERENCES: Bor 1952a; Fairbro<strong>the</strong>rs & Gray 1972; Winter et al. 1982; Barden 1987; Nixon et al.<br />

1987; Hunt & Zaremba 1992; Hor<strong>to</strong>n & Neufeld 1998; Mehr<strong>of</strong>f 2000; Thieret 2003b.<br />

Microstegium vimineum (Trin.) A. Camus, (with long slender shoots, like Salix—willows or<br />

osiers), NEPALESE BROWNTOP, FLEXIBLE SASA GRASS, JAPANESE GRASS, EULALIA, JAPANESE STILT GRASS.<br />

Annual, mat-forming, sometimes in dense colonies; culms creeping at base and rooting at <strong>the</strong><br />

lower nodes, <strong>to</strong> 100(–150) cm long, branched, ascending; ligule a very short membrane <strong>to</strong> ca. 0.8<br />

mm long; leaf blades <strong>to</strong> 2–10(–12) cm long and 3–15 mm wide; inflorescences terminal and lateral<br />

(cleis<strong>to</strong>gamous inflorescences can also be concealed in <strong>the</strong> upper sheaths), <strong>of</strong> (1–)2–5(–6)

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