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1076 POACEAE/STENOTAPHRUM<br />

blades 2–5 mm wide; panicles usually 6–20 cm long, ca. 1/2 as wide as long, <strong>the</strong> lower 1–3 cm <strong>of</strong><br />

each branch bare <strong>of</strong> spikelets; spikelets awnless, 2-flowered (<strong>the</strong> lower floret sterile, <strong>the</strong> upper<br />

floret perfect), glabrous, 1.8–2.4 mm long, sometimes purple-stained, at maturity gaping open<br />

at apex; glumes both present, <strong>the</strong> lower 1/3–1/2 as long as spikelet, acute; upper glume resembling<br />

lemma <strong>of</strong> lower floret; palea <strong>of</strong> <strong>the</strong> lower (neuter) floret inflated, obovate, indurate, <strong>of</strong>ten<br />

apiculate, distinctive, larger than <strong>the</strong> lemma and giving <strong>the</strong> spikelet an expanded, widely gaping<br />

appearance; upper (perfect) floret 1.6–1.9 mm long, <strong>the</strong> lemma and palea firm, but not <strong>to</strong>ugh<br />

and hard nor grain-like. Low areas, moist soils, <strong>of</strong>ten in shade; widespread in e 1/2 <strong>of</strong> TX; se U.S.<br />

from VA s <strong>to</strong> FL w <strong>to</strong> OK and TX, also MO and NM. Apr–Oct, typically early in growing season.<br />

[Panicum hians Elliott]<br />

STENOTAPHRUM Trin. ST. AUGUSTINE GRASS<br />

AA C4 genus <strong>of</strong> 7 species <strong>of</strong> tropical and warm areas <strong>of</strong> <strong>the</strong> world, primarily along <strong>the</strong> Indian<br />

Ocean rim, mainly along seashores or near <strong>the</strong> coast, rarely inland; three species are endemic <strong>to</strong><br />

Madagascar and one is questionably native <strong>to</strong> North America (Clay<strong>to</strong>n & Renvoize 1986; Allred<br />

2003e). It has been suggested that <strong>the</strong> genus is derived from Paspalidium through a progressive<br />

reduction <strong>of</strong> <strong>the</strong> inflorescence branches and expansion <strong>of</strong> <strong>the</strong> central axis <strong>of</strong> <strong>the</strong> inflorescence<br />

(Sauer 1972; Clay<strong>to</strong>n & Renvoize 1986). O<strong>the</strong>r authorities (e.g., Webster 1988), however, disagree<br />

and suggest it is more closely related <strong>to</strong> two Old World genera, Thuarea Pers. and<br />

Uranthoecium Stapf. The swollen inflorescence axis has been postulated <strong>to</strong> be an adaptation <strong>to</strong><br />

dispersal by sea; however, it remains buoyant for only ca. a week (Sauer 1972; Clay<strong>to</strong>n &<br />

Renvoize 1986). Some species are used for pasture, for lawns, or playing fields, or are considered<br />

significant weeds (Watson & Dallwitz 1992). (Greek: steno, narrow and taphros, trench, from<br />

grooves in <strong>the</strong> inflorescence axis in<strong>to</strong> which <strong>the</strong> reduced branches are sunken) (subfamily<br />

Panicoideae, tribe Paniceae)<br />

REFERENCES: Sauer 1972; Crins 1991; Allred 2003e.<br />

Stenotaphrum secundatum (Walter) Kuntze, (with parts arranged along one side), ST. AUGUSTINE<br />

GRASS, CARPET GRASS. S<strong>to</strong>loniferous, sod-forming perennial; culms decumbent or ascending,<br />

rooting at <strong>the</strong> nodes, <strong>the</strong> flowering branches ascending <strong>to</strong> ca. 30 cm tall; leaf sheaths compressed,<br />

keeled, with a few hairs at summit; ligule a minute membrane with a very short ring <strong>of</strong><br />

hairs; leaf blades 5–15(–18) cm long, 5–10 mm wide, � obtuse apically; inflorescences spike-like,<br />

usually 5–10 cm long and 5–10 mm wide, <strong>the</strong> axis wide, flattened, and corky, appearing unbranched<br />

(cryptically a very reduced panicle with short appressed branches—each bearing 1–<br />

3(–5) spikelets—sunken in<strong>to</strong> <strong>the</strong> axis); spikelets 4–5 mm long, awnless, 2-flowered, <strong>the</strong> lower<br />

floret staminate or neuter, <strong>the</strong> upper floret perfect, appressed and sunken in<strong>to</strong> one side <strong>of</strong> <strong>the</strong><br />

axis; lower glume 1–2 mm long, smaller than upper glume, <strong>the</strong> second ca. as long as spikelet;<br />

stamens 3. Commonly planted as a lawn grass, escaping locally <strong>to</strong> a limited extent; tending <strong>to</strong><br />

freeze back in very severe winters, particularly in <strong>the</strong> n part <strong>of</strong> East TX; Bexar, Brazos, Dallas,<br />

Grayson, Harris (BRIT), Colorado, and Travis (Turner et al. 2003) cos.; scattered in TX w <strong>to</strong> e<br />

Cross Timbers and Prairies and Edwards Plateau; se U.S. from VA s <strong>to</strong> FL w <strong>to</strong> MO, OK, and TX,<br />

also CA. Jun–Aug. Native range uncertain; possibly native <strong>to</strong> <strong>the</strong> tropics (e.g., South America—<br />

Webster 1993b) though now widespread. Webster (1988) considered it probably introduced <strong>to</strong><br />

<strong>the</strong> United States. However, it was known along <strong>the</strong> e coast <strong>of</strong> <strong>the</strong> U.S. before 1800 (Sauer 1972),<br />

and recently a number <strong>of</strong> authorities (e.g., Kartesz 1999; Allred 2003e) have considered it native<br />

<strong>to</strong> <strong>the</strong> U.S. This species is widely used as a lawn and turf grass in <strong>the</strong> tropics and subtropics<br />

(Sauer 1972; Crins 1991), and it is also used for soil stabilization (Webster 1988). It can be an aggressively<br />

weedy invader <strong>of</strong> flower beds where grown as a lawn; it can grow vigorously in some<br />

shade. A variegated form is known and is sometimes used as an ornamental in hanging baskets<br />

and greenhouses (Sauer 1972; Allred 2003e). Crins (1991) noted that diploid <strong>plants</strong> have yellow<br />

stigmas and are capable <strong>of</strong> normal sexual reproduction, while triploid (generally used for<br />

lawns) and tetraploid <strong>plants</strong> have purple stigmas and are sterile. ?I

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