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RHYNCHOSPORA/CYPERACEAE<br />

625<br />

34. Culms lax <strong>to</strong> spreading <strong>to</strong> stiffly erect, (10–)20–60(–80) cm tall; leaves<br />

usually 1–2(–3) mm wide; branchlets <strong>of</strong> <strong>the</strong> cymes terminating in small<br />

knobby glomerules, <strong>the</strong> bracts inconspicuous; spikelets (2–)2.5–3(–4)<br />

mm long, roundish, brown <strong>to</strong> dark brown; fertile spikelet scales 1.7–2.3<br />

mm long, <strong>the</strong> midrib usually not extending past tip <strong>of</strong> scale __________ R. globularis<br />

34. Culms stiffly erect, 60–100(–120) cm tall; leaves usually 2–5 mm wide;<br />

branchlets <strong>of</strong> <strong>the</strong> cymes usually terminating in dense glomerules, with<br />

setaceous bracts conspicuous (giving <strong>the</strong> inflorescence a bristly appearance);<br />

spikelets usually 3–4 mm long, elongate, reddish brown; fertile<br />

spikelet scales 2.5–3 mm long, <strong>the</strong> midrib <strong>of</strong>ten extending past tip<br />

<strong>of</strong> scale ___________________________________________________ R. recognita<br />

31. Perianth bristles as long as <strong>the</strong> achenes or longer.<br />

35. Tubercle margins smooth; achenes 1–1.2 mm long ______________________ R. microcarpa<br />

35. Tubercle margins se<strong>to</strong>se (= with minute bristles—use dissecting scope or<br />

strong hand lens); achenes 1.2–1.5 mm long.<br />

36. Achenes nearly round <strong>to</strong> broadly obovoid, <strong>the</strong> surface with horizontal ridges<br />

(<strong>the</strong> cells oblong in outline); tubercle short del<strong>to</strong>id; inflorescences with<br />

spikelet clusters dense; <strong>plants</strong> without evident rhizomes or with short rhizomes<br />

__________________________________________________________ R. caduca<br />

36. Achenes ellipsoid <strong>to</strong> narrowly obovoid, <strong>the</strong> surface without horizontal<br />

ridges, merely minutely honey-combed (<strong>the</strong> cells nearly round in outline);<br />

tubercle del<strong>to</strong>id-subulate (relatively narrower than in R. caduca); inflorescences<br />

with spikelet clusters mostly diffuse; <strong>plants</strong> usually with welldeveloped,<br />

<strong>of</strong>ten elongate rhizomes (<strong>to</strong> 10 cm or more long) ________________ R. mixta<br />

Rhynchospora caduca Elliott, (falling early, short-lived), ANGLE-STEM BEAK SEDGE, ANGLE-STEM<br />

BEAK-RUSH. Perennial 70–150 cm tall, <strong>of</strong>ten with short scaly rhizomes; inflorescences with<br />

spikelet clusters dense; spikelets (3–)4–5 mm long; achenes 1.3–1.5 mm long, nearly round <strong>to</strong><br />

broadly obovoid, lenticular; tubercles 0.5–0.8 mm long, <strong>the</strong> margins with minute bristles.<br />

Ditches, o<strong>the</strong>r low or moist areas; mainly se part <strong>of</strong> East TX and n Gulf Prairies and Marshes,<br />

also Bowie Co. (BRIT) in Red River drainage and Gonzales Co. (Turner et al. 2003) near sw margin<br />

<strong>of</strong> East TX; also e Edwards Plateau; se U.S. from VA s <strong>to</strong> FL w <strong>to</strong> OK and TX. Apr–Nov. [R.<br />

patula A. Gray] This species intergrades with R. mixta (Kral 2002c).<br />

Rhynchospora capitellata (Michx.) Vahl, (having a small head), BROWNISH BEAK SEDGE, POINT-<br />

BEAK BEAK-RUSH. Tufted perennial ca. 20–100 cm tall; inflorescences with clusters <strong>of</strong> spikelets<br />

usually borne singly or in groups <strong>of</strong> ca. 2–3; spikelets 3.5–4(–5) mm long; perianth bristles usually<br />

with retrorse barbs (very rarely antrorse); achenes usually 1.2–1.5 mm long, obovoid, lenticular;<br />

tubercles (0.8–)0.9–1.2(–1.6) mm long. Shorelines, boggy areas; widely scattered in East<br />

TX; se Canada and widespread in <strong>the</strong> e 1/2 <strong>of</strong> <strong>the</strong> U.S., also CA and OR. Jul–Aug. [R. glomerata<br />

(L.) Vahl var. capitellata (Michx.) Kük., R. glomerata var. lep<strong>to</strong>carpa Chapm. ex Brit<strong>to</strong>n, R.<br />

glomerata var. minor Brit<strong>to</strong>n, R. lep<strong>to</strong>carpa (Chapm. ex Brit<strong>to</strong>n) Small] This species intergrades<br />

with R. glomerata (Kral 2002c). Sorrie (2000) indicated that R. lep<strong>to</strong>carpa (Chapm. ex Brit<strong>to</strong>n)<br />

Small (occurring from NC s <strong>to</strong> FL w <strong>to</strong> LA) should be split from R. capitellata; Kral (2002c),<br />

however, noted that <strong>the</strong>re is “a strong overlap” with R. capitellata. Rhynchospora lep<strong>to</strong>carpa was<br />

erroneously mapped for TX by Turner et al. (2003) (see Sorrie 2000).<br />

Rhynchospora cephalantha A. Gray, (bearing heads), BUNCHED BEAK SEDGE. Tufted perennial<br />

0.4–1.1(–1.5) m tall; inflorescences with spikelets in dense, globose or nearly globose, widely<br />

spaced clusters; spikelets 4–5(–6) mm long, each spikelet with only one achene; perianth<br />

bristles usually with retrorse barbs (very rarely antrorse); achenes (1.8–)2–2.3(–2.4) mm long,<br />

obovoid, lenticular; tubercle (1–)1.4–2.4 mm long, triangular-subulate, at l<strong>east</strong> 0.5 mm wide at

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