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keys to the vascular plants of east texas - Botanical Research ...

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474 ARECACEAE/SABAL<br />

REFERENCES: Bailey 1961; Moore 1973; Moore & Uhl 1982; Dahlgren et al. 1985; Dransfield 1986;<br />

Dransfield & Uhl 1986, 1998; Henderson 1986; Uhl & Dransfield 1987, 1999; Zona & Henderson<br />

1989; Tomlinson 1990; Lockett 1991; Henderson et al. 1995; Uhl et al. 1995; Zona 1997, 2000;<br />

Anzizar et al. 1998; Baker et al. 1999; Henderson & Borchsenius 1999; Asmussen et al. 2000.<br />

SABAL Adans. PALMETTO<br />

Perennials with subterranean or aerial, woody or pithy, unbranched spineless stems and a<br />

single apical meristem; leaves long-petioled, <strong>the</strong> blades stiff, evergreen, glabrous, fan-like, longitudinally<br />

pleated <strong>to</strong>ward base, palmately divided over half way (sometimes becoming split <strong>to</strong><br />

base) in<strong>to</strong> many narrowly lanceolate segments; hastula (= flap <strong>of</strong> tissue from petiole where<br />

petiole meets blade) well-developed, adaxial; inflorescence a long panicle with 2 or 3 orders <strong>of</strong><br />

branching, its stalk with leafy bracts consisting <strong>of</strong> a closed, tubular basal sheath and grass-like<br />

short blade; flowers numerous, bisexual, subsessile, small (3.5–6.5 mm long); calyx 3-lobed; petals<br />

3, whitish or greenish, longer than <strong>the</strong> calyx; stamens 6; pistil 1; ovary superior; fruits drupelike,<br />

but with membranous endocarps, black or brownish black.<br />

AA genus <strong>of</strong> 16 species (Zona 2000) <strong>of</strong> trunkless or trunked, unarmed palms ranging from <strong>the</strong><br />

se United States <strong>to</strong> South America. Some are used as a source <strong>of</strong> thatch. Pollination in <strong>the</strong> genus<br />

is mostly by native bees, introduced honeybees, and wasps (Zona 1987, 1990, 2000; Ramp 1989).<br />

Bears are reported <strong>to</strong> disperse <strong>the</strong> seeds <strong>of</strong> Sabal species in Florida (Zona & Henderson 1989).<br />

(Derivation unknown, <strong>the</strong> name possibly derived from an American vernacular name)<br />

REFERENCES: Small 1922; Nixon et al. 1973; Zona 1987, 1990; Lockett & Read 1990; Lockett 1991,<br />

1995, 2003; Ramp & Thien 1995; Goldman 1999.<br />

1. Stems (trunk) aerial; leaves 10–30 per plant, <strong>the</strong> segments 60–115(–145) per leaf, marginally<br />

filiferous (= thread-bearing); hastula 9.5–15.5 cm long, acute <strong>to</strong> acuminate; inflorescences with 3<br />

orders <strong>of</strong> branching (not counting main inflorescence axis); fruits � 13 mm long; species rare in<br />

East TX ________________________________________________________________________ S. mexicana<br />

1. Stems usually subterranean, <strong>the</strong> <strong>plants</strong> thus usually appearing <strong>to</strong> be without a stem (aerial stems<br />

rarely present); leaves 4–10 per plant, <strong>the</strong> segments 15–65(–84) per leaf, marginally not filiferous;<br />

hastula 0.8–4.7 cm long, obtuse; inflorescences usually with 2 orders <strong>of</strong> branching (not counting<br />

main inflorescence axis); fruits � 9 mm long; species widespread and abundant in East TX _________ S. minor<br />

Sabal mexicana Mart., (<strong>of</strong> Mexico), MEXICAN PALMETTO, RIO GRANDE PALMETTO, TEXAS PALMETTO,<br />

PALMA DE MICHAROS. Plant <strong>to</strong> 15 m tall (Lockett [2003] measured a fallen stem ca. 11 m long);<br />

stems (trunk) aerial, 20–35 cm DBH (= diam. br<strong>east</strong> height). Evergreen subtropical woodlands<br />

on deep alluvial soils (TOES 1993), floodplains, riverbanks, and swamps; included based on research<br />

and collections by Lockett (1991, 2003), who worked with <strong>the</strong> diary <strong>of</strong> Fray Isidro Felix<br />

de Espinosa, a priest who accompanied a 1716 Spanish expedition in<strong>to</strong> <strong>the</strong> area that is now San<br />

An<strong>to</strong>nio. Espinosa “reported seeing ‘palmi<strong>to</strong>s legítimos’ at San An<strong>to</strong>nio Springs, source <strong>of</strong> <strong>the</strong><br />

San An<strong>to</strong>nio River, in what is <strong>to</strong>day <strong>the</strong> city <strong>of</strong> San An<strong>to</strong>nio” (Lockett 1991). Lockett subsequently<br />

found S. mexicana in remnant forests near San An<strong>to</strong>nio springs (Bexar Co.). According<br />

<strong>to</strong> Lockett, “Since S. mexicana is commonly cultivated in <strong>the</strong> area <strong>the</strong>re appears <strong>to</strong> be no way <strong>of</strong><br />

knowing, without intensive his<strong>to</strong>rical research, whe<strong>the</strong>r <strong>the</strong> wild specimens are <strong>the</strong> progeny <strong>of</strong><br />

imported trans<strong>plants</strong>, or are descendants <strong>of</strong> a native population, which itself may have been<br />

relocated for landscaping. What we do know is that S. mexicana thrives in <strong>the</strong> area, while S.<br />

minor is lacking.” Zona (2000) also mapped a locality for S. mexicana (county not specified) in<br />

<strong>the</strong> sw part <strong>of</strong> East TX. Lockett and Read (1990) cited <strong>plants</strong> from Jackson and Vic<strong>to</strong>ria cos.<br />

(along Garcitas Creek), and Turner et al. (2003) also mapped Brazos (presumably introduced),<br />

Cameron (native), and Harris (s portion—native status unknown) cos.; probably extreme s portion<br />

<strong>of</strong> East TX, but mainly Rio Grande Valley in <strong>the</strong> South TX Plains and central Gulf Coast in<br />

Gulf Prairies and Marshes (Lockett & Read 1990); Lockett (2003) also noted that <strong>the</strong> species is

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