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keys to the vascular plants of east texas - Botanical Research ...

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910 POACEAE/DIGITARIA<br />

subsessile, <strong>the</strong> o<strong>the</strong>r short-pedicelled), 2-flowered, <strong>the</strong> lower floret staminate or neuter, <strong>the</strong> upper<br />

floret fertile; disarticulation below glumes; lower glume minute or absent; upper glume<br />

well-developed but usually shorter than lemmas; lemma <strong>of</strong> lower floret resembling a glume;<br />

lemma <strong>of</strong> fertile floret with margins thin and flat, not inrolled; stamens 3.<br />

AA C4 genus <strong>of</strong> ca. 200 species <strong>of</strong> tropical <strong>to</strong> warm temperate areas <strong>of</strong> <strong>the</strong> world (Wipff 2003f).<br />

Many species are weedy, a number are important pasture grasses, and some are cultivated for food<br />

(grain) in Africa. As treated here, Digitaria includes Lep<strong>to</strong>loma (e.g., D. cognata), whose inflorescence<br />

appears quite different; see that species for discussion. Three East TX species (D. californica,<br />

D. insularis, and D. patens) were treated in <strong>the</strong> genus Trichachne by Correll and Johns<strong>to</strong>n (1970).<br />

� Digitaria velutina (Forssk.) Beauv., VELVET CARPET GRASS, a federal noxious weed, was erroneously<br />

reported for TX by Kartesz (1999) (Wipff 2003f). (Latin: digitus, finger, from <strong>the</strong> finger-like<br />

arrangement <strong>of</strong> <strong>the</strong> inflorescence branches) (subfamily Panicoideae, tribe Paniceae)<br />

REFERENCES: Henrard 1950; Ebinger 1962; Gould 1963; Veldkamp 1973; Webster & Hatch 1981a,<br />

1981b, 1983, 1990; Webster & Shaw 1982; Webster 1987, 1988; Wipff & Hatch 1988, 1994; Crins<br />

1991; Wipff 2001b, 2003f.<br />

1. Spikelets long-pedicelled (pedicels 2–many times as long as spikelet); inflorescence a muchbranched<br />

and rebranched, open panicle with spikelets far apart ____________________________ D. cognata<br />

1. Spikelets subsessile or on short, appressed pedicels; inflorescence <strong>of</strong> few <strong>to</strong> numerous, unbranched,<br />

spike-like branches (<strong>the</strong>se digitately arranged or along a short axis), <strong>the</strong> spikelets close <strong>to</strong>ge<strong>the</strong>r.<br />

2. Inflorescence branches winged (wings <strong>of</strong>ten as wide as central part <strong>of</strong> branch); spikelets glabrous<br />

<strong>to</strong> short-pubescent.<br />

3. Spikelets 1.2–2.3 mm long; lemma <strong>of</strong> fertile floret dark brown <strong>to</strong> purplish black at maturity.<br />

4. Spikelets (1.7–)1.9–2.3 mm long ___________________________________________ D. ischaemum<br />

4. Spikelets 1.2–1.7 mm long _________________________________________________ D. violascens<br />

3. Spikelets 2.2–3.7 mm long; lemma <strong>of</strong> fertile floret light brown or grayish.<br />

5. Lemma <strong>of</strong> lower floret (lemma appearing glume-like because <strong>of</strong> minute or absent lower<br />

glume) <strong>of</strong> <strong>the</strong> nearly sessile spikelets with <strong>the</strong> 7 (sometimes apparently 5) veins � equally<br />

spaced, <strong>of</strong>ten but not always densely villous marginally (note that spikelets are in pairs,<br />

one pedicelled and one nearly sessile); lower glume <strong>of</strong> nearly sessile spikelets rounded<br />

<strong>to</strong> truncate or bifid, usually 0.1–0.3 mm long _____________________________________ D. bicornis<br />

5. Lemma <strong>of</strong> lower floret <strong>of</strong> <strong>the</strong> nearly sessile spikelets with <strong>the</strong> 7 (sometimes apparently<br />

5) veins not equally spaced, <strong>the</strong> lateral ones crowded at margins, glabrous or inconspicuously<br />

pubescent marginally; lower glume <strong>of</strong> nearly sessile spikelets obtuse or acute,<br />

<strong>of</strong>ten more than 0.3 mm long.<br />

6. Spikelets 2.2–3.3 mm long; upper glume 0.8–1.8 mm long; leaf blades usually densely<br />

pubescent, usually on both surfaces; lemma <strong>of</strong> lower floret minutely scabrous on<br />

lateral veins (as seen under a dissecting scope) _____________________________ D. sanguinalis<br />

6. Spikelets 2.8–4.1 mm long; upper glume 1.5–2.7 mm long; leaf blades glabrous or<br />

sparsely pubescent, any hairs usually on upper surface only; lemma <strong>of</strong> lower floret<br />

smooth on lateral veins _____________________________________________________ D. ciliaris<br />

2. Inflorescence branches not winged or only minutely so; spikelets silky-pubescent with long<br />

white-, tan-, buff-, or purplish-silky hairs OR not so.<br />

7. Spikelets silky-pubescent, <strong>the</strong> inflorescences appearing conspicuously white-, tan-, buff-,<br />

or purplish-silky-pubescent at arm’s length; spikelets 3–5.9 mm long.<br />

8. Back <strong>of</strong> lemma <strong>of</strong> lower, sterile floret usually with pubescence or apparently so (in addition<br />

<strong>to</strong> being conspicuously hirsute marginally); inflorescence usually tan- or buff-tinged<br />

due <strong>to</strong> conspicuous pubescence on spikelets ____________________________________ D. insularis<br />

8. Back (outer surface) <strong>of</strong> lemma <strong>of</strong> lower, sterile floret (lemma glume-like because <strong>of</strong> minute<br />

1st glume) glabrous at l<strong>east</strong> in a broad lengthwise band down <strong>the</strong> middle (but marginally<br />

densely hirsute); inflorescence usually white- or purplish-tinged due <strong>to</strong> conspicuous<br />

pubescence on spikelets.

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