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keys to the vascular plants of east texas - Botanical Research ...

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POACEAE 797<br />

Dissection is usually necessary <strong>to</strong> see <strong>the</strong> parts <strong>of</strong> <strong>the</strong> grass spikelet, and because <strong>of</strong> <strong>the</strong> small<br />

size <strong>of</strong> <strong>the</strong> structures involved (e.g., lemmas, paleas), a dissecting scope or at minimum a hand<br />

lens is <strong>of</strong>ten necessary for definitive identification. In <strong>the</strong> <strong>keys</strong>, measurements <strong>of</strong> glumes, lemmas,<br />

or spikelets do not include awns (= hair-like or bristle-like appendages) if <strong>the</strong>se are<br />

present. (subclass Commelinidae—Cronquist; order Poales—APG II)<br />

FAMILY RECOGNITION IN THE FIELD: mostly herbs with 2-ranked leaves having sheathing bases,<br />

free blades, and ligules; culms round, with hollow or solid internodes; flowers small, inconspicuous,<br />

reduced <strong>to</strong> stamens and pistils, subtended by 2 scale-like bracts each, and arranged in<br />

very reduced spikes called spikelets; fruit a caryopsis; <strong>the</strong> � similar Cyperaceae (SEDGES) have<br />

3-ranked leaves, <strong>of</strong>ten 3-sided culms with solid internodes, each flower subtended by 1 scalelike<br />

bract, and fruit an achene; <strong>the</strong> � similar Juncaceae (RUSHES) have flowers with a small 6parted<br />

perianth and capsular fruit.<br />

REFERENCES: Nash 1909, 1912; Nash & Hitchcock 1915; Hitchcock 1931–1939, 1935, 1951; Silveus<br />

1933; Tharp 1952a; Shinners 1954; Gould 1957b, 1968a, 1968b, 1973, 1975b, 1978; Box & Gould<br />

1959; deWet 1981; Stebbins 1981a, 1982, 1985; Estes et al. 1982; Gould & Shaw 1983; McVaugh<br />

1983; Campbell 1985; Coughenour 1985; Dahlgren et al. 1985; Redmann 1985; Clay<strong>to</strong>n &<br />

Renvoize 1986; Soderstrom et al. 1987; Hignight et al. 1988; Tucker 1988, 1990, 1996; Crins 1991;<br />

Allen 1992b, 2004a; Watson & Dallwitz 1992; Powell 1994; Barker et al. 1995, 1998[1999]; Clark et<br />

al. 1995; Kellogg & Linder 1995; Kellogg et al. 1996; Petersen & Seberg 1997; Peterson et al. 1997;<br />

Hsaio et al. 1998; Kucera 1998; Soreng & Davis 1998, 2000; Hatch et al. 1999; Grass Phylogeny<br />

Working Group 2000, 2001; Hilu 2000; Hilu & Alice 2000, 2001; Jacobs & Everett 2000;<br />

Judziewicz et al. 2000; Kellogg 2000a, 2000b; Spangler 2000; Zuloaga et al. 2000; Mohlenbrock<br />

2001; Hatch 2002; Ma<strong>the</strong>ws et al. 2002; Soreng et al. 2003; Zuloaga et al. 2003; C<strong>of</strong>fey & Stevens<br />

2004; Sánchez-Moreiras et al. 2004.<br />

KEY TO GENERA OF POACEAE<br />

1. Culms woody, persisting for more than 1 season (bamboos).<br />

2. Culms terete (= rounded); upper nodes <strong>of</strong>ten puberulent; base <strong>of</strong> midvein on lower leaf surface<br />

without short hairs; branches 3–6 per node; <strong>plants</strong> <strong>of</strong> moist woods or low areas, native in<br />

e part <strong>of</strong> East TX _______________________________________________________________ Arundinaria<br />

2. Culms flattened above node on 1 side; upper nodes glabrous; base <strong>of</strong> midvein on lower leaf<br />

surface <strong>of</strong>ten with short hairs along one side; branches usually 2(–3) per node; <strong>plants</strong> introduced<br />

in East TX, persisting and spreading from cultivation ___________________________ Phyllostachys<br />

1. Culms not woody, lasting 1 season only (sometimes lasting 2 seasons and somewhat woody in<br />

Arundo).<br />

3. Plants typically 2–7 m tall, <strong>of</strong>ten reed-like; inflorescences typically large, sometimes plumose<br />

panicles up <strong>to</strong> 60(–130) cm long______________________________________________ Key A (page /798)<br />

3. Plants usually � 2(–3+) m tall, not reed-like; inflorescences various.<br />

4. Leaves with blades less than 1(–1.5) cm long, in clusters; inflorescences very inconspicuous<br />

and obscured by <strong>the</strong> leaves, typically reduced <strong>to</strong> a single(–few) spikelet ________ Monanthochloe<br />

4. Leaves with blades more than 1.5 cm long, usually not in clusters; inflorescences usually<br />

conspicuous.<br />

5. Spikelets fused with or closely fitted in<strong>to</strong> <strong>the</strong> axis <strong>of</strong> <strong>the</strong> inflorescence or inflorescence<br />

branches, forming a solid cylindrical or flattened spike ________________________ Key B (page /799)<br />

5. Spikelets nei<strong>the</strong>r fused with nor closely fitted in<strong>to</strong> <strong>the</strong> axis <strong>of</strong> <strong>the</strong> inflorescence or inflorescence<br />

branches (but may be sessile and appressed).<br />

6. Spikelets unisexual, <strong>the</strong> staminate and pistillate spikelets conspicuously different <strong>to</strong><br />

<strong>the</strong> naked eye _____________________________________________________ Key C (page /799)<br />

6. Spikelets perfect, or if unisexual, not conspicuously different <strong>to</strong> <strong>the</strong> naked eye.<br />

7. Spikelets with 2–many perfect florets and thus usually 2–many fruits forming per<br />

spikelet.

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