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keys to the vascular plants of east texas - Botanical Research ...

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454 AMARYLLIDACEAE/LEUCOJUM<br />

1. Plants flowering summer–early fall (Jul–Sep); leaves wi<strong>the</strong>ring before flowering time OR leaves<br />

fresh at flowering time, distinctly wider above middle before tapering <strong>to</strong>ward end, narrowly<br />

oblanceolate in overall shape, 2–6 cm wide, � not coriaceous; perianth tube 7–13.5 cm long;<br />

corona (2.5–)3.3–4(–4.5) cm long; free portion <strong>of</strong> filaments 2.5–4 cm long; <strong>plants</strong> <strong>of</strong> variable, but<br />

sometimes mesic habitats _______________________________________________________ H. occidentalis<br />

Hymenocallis liriosme (Raf.) Shinners, (lily-smell or fragrant lily), SPRING SPIDER-LILY, WESTERN<br />

SPIDER-LILY, FRAGRANT SPIDER-LILY, WESTERN MARSH SPIDER-LILY, LOUISIANA MARSH SPIDER-LILY.<br />

Plant <strong>to</strong> 68(–88) cm tall; leaves shiny green; flowers (3–)5–12; perianth similar <strong>to</strong> that <strong>of</strong> H.<br />

occidentalis but slightly smaller, <strong>the</strong> tepals 6–9 cm long, 8 mm or less wide. Stream bot<strong>to</strong>ms,<br />

ditches, pond margins, o<strong>the</strong>r wet areas, sometimes in shallow water; widespread in Pineywoods<br />

and e Post Oak Savannah; also n Gulf Prairies and Marshes; AL, AR, LA, MS, OK, and TX. Mar–<br />

May. [H. caroliniana <strong>of</strong> TX authors, not (L.) Herb., H. galves<strong>to</strong>nensis (Herb.) Baker, H. galves<strong>to</strong>nensis<br />

subsp. angustifolia Traub, Pancratium liriosme Raf.] Shinners (1951a) correctly recognized<br />

<strong>the</strong>re was only one spring-flowering SPIDER-LILY in TX; unfortunately, later authors (e.g., Correll<br />

& Johns<strong>to</strong>n 1970; Jones et al. 1997; Diggs et al. 1999; Turner et al. 2003) thought <strong>the</strong>re were two<br />

species and called one <strong>of</strong> <strong>the</strong>m H. caroliniana, a species whose type specimen is actually <strong>of</strong> a<br />

different species and even genus, Pancratium maritimum L. (Smith & Flory 2002). m/289<br />

Hymenocallis occidentalis (J. Le Conte) Kunth, (western), WOODLAND SPIDER-LILY, HAMMOCK SPIDER-<br />

LILY, NORTHERN SPIDER-LILY, SUMMER SPIDER-LILY. Plant similar <strong>to</strong> H. liriosme, (40–)50–75 cm tall;<br />

leaves glaucous when young; flowers 3–9; perianth extremely large and showy, <strong>the</strong> tepals white,<br />

green-striped on keel, (7–)8.5–11.5 cm long, 10 mm or less wide, <strong>the</strong> corona white with small yellowish<br />

green eye.<br />

1. Leaves wi<strong>the</strong>ring before flowering time _________________________________________________ var. eulae<br />

1. Leaves fresh at flowering time ___________________________________________________ var. occidentalis<br />

var. eulae (Shinners) Ger. L. Sm. & Flory, (for Eula Whitehouse, 1892–1974, herbarium botanist<br />

and collec<strong>to</strong>r at Sou<strong>the</strong>rn Methodist University, and author/artist <strong>of</strong> Texas Flowers in Natural<br />

Colors (1936), <strong>the</strong> first color-illustrated guide <strong>to</strong> TX wildflowers). Leaves appearing in late winter,<br />

wi<strong>the</strong>ring in late spring before flowering, suberect; scape bracts 4–5 cm long. Sandy pine or<br />

mixed hardwood forests, in heavy soils near streams, seepage slopes; Shelby, Smith, Tyler (BRIT),<br />

Burleson (TAMU), Madison (Neill & Wilson 2001), Anderson, Gregg, Grimes, Panola, and Red<br />

River (Turner et al. 2003) cos.; OK and TX. Summer–early fall. [H. eulae Shinners] Smith and<br />

Flory (2002) indicated that this taxon is in need <strong>of</strong> additional study. m/289<br />

var. occidentalis. Leaves appearing in late winter, fresh at flowering and subsequently wi<strong>the</strong>ring,<br />

arching outward; scape bracts 4.5–7 cm long. Floodplain forests, wooded hillsides, sometimes<br />

in upland mesic forests; according <strong>to</strong> Smith and Flory (2002) and Smith and Garland<br />

(2003), this taxon occurs widely in <strong>the</strong> e U.S. from NC s <strong>to</strong> FL w <strong>to</strong> IL, AR, and LA but not in TX.<br />

Turner et al. (2003) mapped H. caroliniana (a misapplied name) widely in East TX and <strong>the</strong> Gulf<br />

Prairies and Marshes. It is unclear whe<strong>the</strong>r <strong>the</strong>y were referring <strong>to</strong> H. liriosome or possibly <strong>to</strong> H.<br />

occidentalis var. occidentalis. However, we have not seen any specimens that are unambiguously<br />

var. occidentalis and are thus only provisionally including this species for East TX; no distribution<br />

map is provided. Summer–early fall. [H. caroliniana <strong>of</strong> authors, not (L.) Herb.] This<br />

taxon has long been cultivated in sou<strong>the</strong>rn gardens, and specimens “<strong>of</strong>ten have been identified<br />

as H. caroliniana (Linnaeus) Herbert on herbarium sheets and in floras” (Smith & Flory 2002).<br />

However, that name is now known <strong>to</strong> be a synonym <strong>of</strong> a different species (Smith & Flory 2002).<br />

LEUCOJUM L. SNOWFLAKE<br />

AA genus <strong>of</strong> ca. 10 species (Straley & Utech 2002c) ranging from Europe <strong>to</strong> nw Africa and sw<br />

Asia; related <strong>to</strong> and sometimes confused with Galanthus (Meerow et al. 1999 [2000], 2000b).<br />

However, Leucojum has “perianth segments that are all equal in size, and hollow stems that are

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