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972 POACEAE/LIMNODEA<br />

swamps; Bowie, Harris, and Liberty (SBSC) cos.; also Johnson Co. (Carr 17715, BRIT) just <strong>to</strong> <strong>the</strong> w<br />

<strong>of</strong> East TX, and Fort Bend Co. (SBSC) in <strong>the</strong> n part <strong>of</strong> Gulf Prairies and Marshes; Snow (1997,<br />

2003a) cited no TX specimens; e U.S. from IN s <strong>to</strong> FL w <strong>to</strong> MO and TX. Spring–fall. [Diplachne<br />

panicoides (J. Presl) McNeill] Considered by Gould (1975b) <strong>to</strong> be introduced from Brazil, but<br />

treated more recently as a U.S. native (e.g., Kartesz 1999; Snow 2003a).<br />

Lep<strong>to</strong>chloa virgata (L.) Beauv., (wand-like), TROPIC SPRANGLETOP. Tufted perennial; culms 30–<br />

200 cm tall, usually erect; ligule a membrane 0.2–1 mm long; axils <strong>of</strong> inflorescence branches<br />

pilose; lemmas awnless or with awns <strong>to</strong> 3(–11) mm long. Ditches or o<strong>the</strong>r moist sites; Bexar<br />

(Snow 1997) and Jefferson (Turner et al. 2003) cos. at extreme s margin <strong>of</strong> East TX; mainly Gulf<br />

Prairies and Marshes; FL, SC, and TX. May–Nov. [Cynosurus virgatus L., L. domingensis (Jacq.)<br />

Trin.] While Nicora (1995) divided this species in<strong>to</strong> several taxa, Snow (1997) indicated that<br />

though it is highly polymorphic, separation in<strong>to</strong> infraspecific taxa is not warranted. Snow<br />

(2003a) also noted that “Awn length and lemma pubescence vary continuously and independently,<br />

precluding <strong>the</strong>ir use in recognizing additional taxa.”<br />

LIMNODEA L.H. Dewey OZARK GRASS<br />

AA monotypic C3 genus <strong>of</strong> <strong>the</strong> s United States (Snow ined.). It is related <strong>to</strong> <strong>the</strong> perennial genus<br />

Cinna and is sometimes treated in that genus (Tucker 1996; Yatskievych 1999). However,<br />

Brandenburg and Thieret (2000) enumerated numerous differences between <strong>the</strong>se genera (e.g.,<br />

Limnodea is an annual, is smaller, and has longer lemma awns) and concluded that <strong>the</strong>y are<br />

not congeneric. (Name altered from Limnas, a related Old World genus) (subfamily Pooideae,<br />

tribe Poeae)<br />

REFERENCES: Tucker 1996; Brandenburg & Thieret 2000; Snow ined.<br />

Limnodea arkansana (Nutt.) L.H. Dewey, (<strong>of</strong> Arkansas), OZARK GRASS. Annual 15–60(–100) cm<br />

tall; ligule a lacerate-ciliate membrane 1–2 mm long; leaf blades 2–8 mm wide, glabrous or<br />

more <strong>of</strong>ten hispidulous or hispid on upper and lower surfaces; inflorescence a narrow but not<br />

cylindrical panicle, erect or nodding at tip; spikelets 1-flowered, 3–4 mm long excluding awn,<br />

disarticulating below <strong>the</strong> glumes; lemma usually slightly 2-<strong>to</strong>o<strong>the</strong>d at apex, awned from back<br />

at or near <strong>the</strong> base <strong>of</strong> <strong>the</strong> 2 teeth, <strong>the</strong> awn (4–)8–12(–14.5) mm long, geniculate; an<strong>the</strong>rs 3. Prairies<br />

and disturbed areas, calcareous soils, <strong>of</strong>ten in sand; throughout most <strong>of</strong> TX except extreme<br />

w and nw; se U.S. from SC s <strong>to</strong> FL w <strong>to</strong> OK and TX, except GA. Mar–Jun. [Cinna arkansana<br />

(Nutt.) G.C. Tucker]<br />

LOLIUM L. RYE GRASS, DARNEL<br />

Annuals or perennials; leaves auricled; ligule a membrane; inflorescence an unbranched spike<br />

(rarely branched) with sessile spikelets in 2 ranks on opposite sides <strong>of</strong> <strong>the</strong> axis, <strong>the</strong> axis remaining<br />

intact, disarticulation occurring above glume and between florets; spikelets 1 per node,<br />

placed with edge (keels <strong>of</strong> lemmas) against axis <strong>of</strong> inflorescence, sometimes partially in concavity<br />

in axis, without a glume on that side, (2–)5–15(–22)-flowered.<br />

AA temperate Eurasian C3 genus <strong>of</strong> 5–8 diploid species (Bulinska-Radomska & Lester 1988;<br />

Loos 1993a; Terrell ined.), including valuable fodder and lawn grasses as well as significant<br />

weeds. Lolium is considered <strong>to</strong> be <strong>of</strong> recent origin, and <strong>the</strong> species are closely related (Charmet<br />

et al. 1997). While previously placed in <strong>the</strong> tribe Triticeae based on its spicate inflorescence,<br />

Lolium is now recognized as being in <strong>the</strong> Poeae. A number <strong>of</strong> workers (e.g., Bulinska-Radomska<br />

& Lester 1988) have noted similarities between <strong>the</strong> genera Lolium, Festuca, and Vulpia, and<br />

hybrids between Lolium and Festuca are known (Watson & Dallwitz 1992; Terrell ined.).<br />

Darbyshire (1993) suggested that Festuca subgenus Schedonorus, <strong>the</strong> broad-leaved fescues (including<br />

F. arundinacea) be shifted <strong>to</strong> Lolium. However, based on recent molecular studies (e.g.,

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