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keys to the vascular plants of east texas - Botanical Research ...

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1004 POACEAE/PAPPOPHORUM<br />

blades broad, 2–7 cm long, 5–22 mm wide, only 4–6 times as long as wide; inflorescence open;<br />

pedicels elongate, 9–20 mm long; spikelets 1–1.4 mm long, with sparse pubescence. Moist<br />

weedy areas; Travis Co. (Turner et al. 2003) near w margin <strong>of</strong> East TX; also Cameron Co. (Gould<br />

1975b; Turner et al. 2003) at <strong>the</strong> s tip <strong>of</strong> TX; in <strong>the</strong> U.S. known only from TX. Aug–Oct. Native <strong>to</strong><br />

Latin America. According <strong>to</strong> Gould (1975b), this species is “probably not established” in TX. I<br />

Panicum verrucosum Muhl., (warty), WARTY PANICUM, WARTY PANIC GRASS. Glabrous annual with<br />

erect <strong>to</strong> decumbent culms <strong>to</strong> 120(–150) cm long; ligule 0.5 mm or less long; leaf blades 3–10 mm<br />

wide; panicle few-flowered; spikelets 1.7–2.2(–2.5) mm long, conspicuously warty, glabrous.<br />

Moist <strong>to</strong> marshy areas and open woodlands, sandy soils; widespread in Pineywoods and Post<br />

Oak Savannah; e U.S. from MA s <strong>to</strong> FL w <strong>to</strong> MO and TX. Jul–Nov. See description <strong>of</strong> P.<br />

brachyanthum for comparison with that similar species.<br />

Panicum virgatum L., (twiggy, wand-like), SWITCH GRASS. Large rhizoma<strong>to</strong>us perennial 0.6–2(–3)<br />

m tall; nodes glabrous; leaf sheaths usually glabrous; ligule a fringed membrane (1.5–)2–6 mm<br />

long; leaf blades 3–15 mm wide, glabrous or <strong>of</strong>ten pilose at base; panicle large, usually 15–55 cm<br />

long, <strong>the</strong> branches spreading <strong>to</strong> ascending; spikelets acuminate-pointed, 2.8–5 mm long, usually<br />

gaping open at apex; lower glume � 1/2 as long as spikelet. Low moist areas and prairies;<br />

throughout TX; se Canada and throughout most <strong>of</strong> <strong>the</strong> U.S. Aug–Nov. SWITCH GRASS was one <strong>of</strong><br />

<strong>the</strong> dominants in <strong>the</strong> original tall grass prairie; it is considered one <strong>of</strong> <strong>the</strong> “big four” tall grasses<br />

along with Andropogon gerardii, Schizachyrium scoparium, and Sorghastrum nutans. However,<br />

its abundance in native grasslands decreases with grazing (Freckmann & Lelong 2003b). This<br />

species is seen in East TX in two growth forms: 1) LOWLAND SWITCH GRASS—very large isolated<br />

clumps <strong>of</strong>ten nearly 2(–3) m tall, usually in low moist areas, and 2) UPLAND SWITCH GRASS—<br />

shorter <strong>plants</strong> not apparently clumped, with culms more scattered along <strong>the</strong> creeping rhizomes,<br />

found in drier sites. Davis et al. (1995) pointed out additional differences and suggested<br />

<strong>the</strong> variation is possibly worthy <strong>of</strong> taxonomic recognition. The morphological variation seen is<br />

not surprising since <strong>the</strong> species is quite variable in terms <strong>of</strong> chromosomal makeup—“<strong>plants</strong> in a<br />

small area can range from diploid through duodecaploid [12X], with dysploid derivatives”<br />

(Freckmann & Lelong 2003b). While native <strong>to</strong> North America, this species has been introduced<br />

as a forage species <strong>to</strong> o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> world (Freckmann & Lelong 2003b).<br />

PAPPOPHORUM Schreb. PAPPUS GRASS<br />

AA New World C4 genus <strong>of</strong> 8 species (Reeder & Toolin 1989; Reeder 2003) ranging from <strong>the</strong> s<br />

U.S. <strong>to</strong> Argentina. There has been taxonomic confusion in <strong>the</strong> past, with <strong>the</strong> number <strong>of</strong> species<br />

recognized ranging from 8–20 (Reeder & Toolin 1989). Preliminary molecular data (Hilu & Alice<br />

2001) suggest an affinity with some species <strong>of</strong> Eragrostis. Salt glands, <strong>to</strong> dispose <strong>of</strong> excess<br />

salt, have been observed in two species (as in a number <strong>of</strong> o<strong>the</strong>r species <strong>of</strong> subfamily<br />

Chloridoideae) (Taleisnik & An<strong>to</strong>n 1988). (Greek: pappos, pappus, and phoros, bearing, in reference<br />

<strong>to</strong> <strong>the</strong> pappus-like crown <strong>of</strong> <strong>the</strong> lemma) (subfamily Chloridoideae, tribe Pappophoreae)<br />

REFERENCES: Pensiero 1986; Taleisnik & An<strong>to</strong>n 1988; Reeder & Toolin 1989; Reeder 2003.<br />

Pappophorum bicolor E. Fourn., (two-colored), PINK PAPPUS GRASS. Tufted perennial <strong>to</strong> 100 cm<br />

tall; ligule a ring <strong>of</strong> hairs ca. 1 mm long; leaf blades 5 mm or less wide; inflorescence a contracted<br />

spike-like panicle <strong>to</strong> ca. 20 cm long, <strong>of</strong>ten with some erect-spreading branches, pink or<br />

purplish at maturity; spikelets 6–8(–10) mm long (including numerous awns), composed <strong>of</strong> 2–3<br />

perfect florets with 1–2 reduced florets above, disarticulating above <strong>the</strong> glumes, <strong>the</strong> florets falling<br />

as a unit; lemmas rounded on <strong>the</strong> back, indistinctly many-veined, with 11–15 awns; bodies<br />

<strong>of</strong> lower lemmas 2.5–4 mm long; awns variable in length, <strong>the</strong> longest 2.5–5(–6) mm long, <strong>to</strong> 1.5<br />

times <strong>the</strong> length <strong>of</strong> <strong>the</strong> lemma body; reduced florets similar <strong>to</strong> fertile ones but smaller. Grassy<br />

or brushy areas, roadsides, and along streams; Bexar and DeWitt (TAES) cos. near sw margin <strong>of</strong><br />

East TX; mainly s and w TX; in <strong>the</strong> U.S. known only from TX (Reeder 2003); also n Mexico.<br />

Apr–Nov.

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