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keys to the vascular plants of east texas - Botanical Research ...

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MUHLENBERGIA/POACEAE<br />

983<br />

10. Lemma with a tuft <strong>of</strong> hairs at base; culms glabrous and shining between <strong>the</strong> nodes<br />

and at <strong>the</strong> nodes OR puberulent and dull just below <strong>the</strong> nodes.<br />

11. Lemma with a conspicuous awn (3–)5–10(–18) mm long; culms puberulent and<br />

dull just below <strong>the</strong> nodes (use hand lens or dissecting scope); species rare, if<br />

present, in East TX __________________________________________________ M. sylvatica<br />

11. Lemma awnless or with an awn <strong>to</strong> 3 mm long (rarely <strong>to</strong> 8 mm in M. bushii);<br />

culms puberulent and dull just below <strong>the</strong> nodes OR nodes and internodes glabrous<br />

and shiny; including species widely scattered in East TX.<br />

12. Glumes overlapping nearly <strong>to</strong> middle, ovate, abruptly tapering <strong>to</strong> an awn<br />

tip; <strong>plants</strong> with few axillary panicles, but those present usually long-exserted,<br />

<strong>the</strong> peduncles <strong>to</strong> 11 cm long _____________________________________ M. sobolifera<br />

12. Glumes not overlapping or overlapping only at base, lanceolate, gradually<br />

tapering <strong>to</strong> an awn tip; <strong>plants</strong> with numerous axillary panicles, <strong>the</strong>se not<br />

long-exserted (<strong>of</strong>ten partially enclosed in leaf sheath), <strong>the</strong> peduncles usually<br />

only 1–2 cm long.<br />

13. Glumes 1/2–2/3 as long as lemma, 1.2–2(–2.5) mm long; ligule 0.7 mm<br />

or less long; terminal panicle <strong>of</strong>ten well-exserted; leaves <strong>of</strong> <strong>the</strong> lateral<br />

branches <strong>of</strong>ten shorter and narrower than those <strong>of</strong> <strong>the</strong> main culms _______ M. bushii<br />

13. Glumes 3/4 as long <strong>to</strong> slightly longer than lemma, usually ca. 2–4.1 mm<br />

long; ligule 0.7–1.4(–1.7) mm long; terminal panicle usually not wellexserted<br />

(at l<strong>east</strong> base enclosed in subtending leaf sheath); leaves <strong>of</strong><br />

<strong>the</strong> lateral branches about same length and width as those <strong>of</strong> <strong>the</strong> main<br />

culms _____________________________________________________ M. frondosa<br />

Muhlenbergia bushii R.W. Pohl, (for Benjamin Franklin Bush, 1858–1937, amateur botanist <strong>of</strong><br />

Missouri, and discoverer <strong>of</strong> this species), NODDING MUHLY. Rhizoma<strong>to</strong>us perennial; culms 30–<br />

100 cm tall, erect, becoming much-branched above, nodes and internodes glabrous and shiny;<br />

leaf sheaths rounded or slightly angled; ligule 0.7 mm or less long; panicles numerous, slender;<br />

spikelets 2.5–3.6 mm long; lemma pilose basally, awnless or with awn rarely <strong>to</strong> 8 mm long. Rich<br />

or low woods, along streams, <strong>of</strong>ten calcareous conditions; Grayson, Hunt, McLennan, Van Zandt<br />

(BRIT), Dallas, Navarro (TAES), and Travis (Turner et al. 2003) cos.; also Cross Timbers and Prairies;<br />

much <strong>of</strong> <strong>the</strong> c U.S. from IN s <strong>to</strong> LA w <strong>to</strong> NE and TX, also GA and WV. Jul–Oct. [M.<br />

brachyphylla Bush]<br />

Muhlenbergia capillaris (Lam.) Trin., (hair-like), HAIRY-AWN MUHLY, LONG-AWN HAIR GRASS, SLEN-<br />

DER MUHLY, HAIR GRASS. Densely tufted perennial without rhizomes; culms erect, 60–100(–150)<br />

cm tall; leaf sheaths rounded, <strong>the</strong> basal ones <strong>of</strong>ten fibrous but not extensively so; ligule 1.8–8<br />

(–10) mm long; panicles open, diffuse, 15–40(–60) cm long, 5–30(–40) cm wide, <strong>the</strong> branches<br />

and pedicels capillary and widely spreading at maturity; spikelets usually (2.6–)3–4.5(–5) mm<br />

long; glume body � 1/2 as long as lemma body; upper glume usually with awn 0.5–5 mm long;<br />

lemma occasionally with setaceous teeth <strong>to</strong> ca. 2 mm long, with variable awn (0–)2–15(–18)<br />

mm long. Sandy or rocky forests, openings; widespread in East TX; also Gulf Prairies and<br />

Marshes, scattered in <strong>the</strong> Cross Timbers and Prairies, and w <strong>to</strong> Taylor Co. (Peterson 2003b;<br />

Turner et al. 2003); e U.S. from MA s <strong>to</strong> FL w <strong>to</strong> KS and TX. Aug–Nov. This species is grown as an<br />

ornamental (Peterson 2003b).<br />

Muhlenbergia dubia E. Fourn., (doubtful), PINE MUHLY, PINELAND MUHLY. Densely tufted perennial<br />

without rhizomes; culms erect, 30–100 cm tall; leaf sheaths rounded; ligule 4–10 mm long;<br />

panicle 10–40 cm long, <strong>to</strong> 2.4 cm wide, contracted, densely-flowered, <strong>the</strong> branches with flowers<br />

<strong>to</strong> base; spikelets (3.8–)4–5 mm long; lemma glabrous proximally, scabrous distally, awnless or<br />

with awn <strong>to</strong> 6 mm long. Slopes, rock outcrops; Travis Co. (M.C. Johns<strong>to</strong>n 12442 (TAES, TEX);<br />

Denny 2003) near w margin <strong>of</strong> East TX; since this species was added after map pages for <strong>the</strong><br />

flora were completed, no county distribution map is provided; mainly Trans-Pecos but also

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