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keys to the vascular plants of east texas - Botanical Research ...

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624 CYPERACEAE/RHYNCHOSPORA<br />

17. Surface <strong>of</strong> achenes horizontally ridged, rugose (= wrinkled), pitted, or<br />

honeycombed-reticulate (use magnification).<br />

22. Perianth bristles absent (can be absent in R. perplexa); achenes 0.9<br />

mm or less long; tubercle minute, <strong>to</strong> 0.15 mm long.<br />

23. Surface <strong>of</strong> achenes horizontally ridged (use hand lens or dissecting<br />

scope), not glassy in appearance ______________________ R. pusilla<br />

23. Surface <strong>of</strong> achenes faintly reticulate (using dissecting scope),<br />

but not horizontally ridged, somewhat glassy in appearance<br />

____________________________________________________ R. divergens<br />

22. Perianth bristles usually present; achenes 1–2.2 mm long; tubercle<br />

larger, > 0.15 mm long.<br />

24. Culms and leaves filiform-wiry, almost thread-like, 1 mm or less wide.<br />

25. Perianth bristles shorter than achenes; tubercle del<strong>to</strong>id <strong>to</strong><br />

del<strong>to</strong>id-conical, 0.3–0.6 mm long; spikelets 3–4(–4.5) mm<br />

long ______________________________________________ R. rariflora<br />

25. Perianth bristles longer than achenes; tubercle taller and<br />

narrower, del<strong>to</strong>id-subulate, 0.8–1.5 mm long; spikelets 4–5<br />

mm long _______________________________________ R. stenophylla<br />

24. Culms and leaves s<strong>to</strong>uter, not filiform-wiry, usually > 1 mm wide.<br />

26. Faces <strong>of</strong> <strong>the</strong> achenes flat or concave.<br />

27. Spikelets 5–7(–9) mm long; achenes 2 times as long as<br />

wide or longer, 2–2.5 mm long; tubercle 0.8–1.2 mm<br />

long __________________________________________ R. inexpansa<br />

27. Spikelets 2–3.5 mm long; achenes < 2 times as long as<br />

wide, 1–1.3 mm long; tubercle 0.2–0.5(–0.7) mm long.<br />

28. Perianth bristles 6, surpassing <strong>the</strong> tubercle, <strong>of</strong>ten<br />

widely spreading; leaf blades 3–6 mm wide _________ R. elliottii<br />

28. Perianth bristles 0–3(–6), absent or rudimentary, if<br />

present, < three-fourths as long as <strong>the</strong> achenes, not<br />

widely spreading; leaf blades 1.5–2.5 mm wide _____ R. perplexa<br />

26. Faces <strong>of</strong> <strong>the</strong> achenes convex or swollen, sometimes with a central bulge.<br />

29. Inflorescence branches spreading nearly at right angles <strong>to</strong> <strong>the</strong> main inflorescence axis, <strong>the</strong><br />

inflorescence appearing very open, each spikelet or small cluster <strong>of</strong> spikelets on a slender<br />

stalk; achenes yellowish <strong>to</strong> greenish yellow, tan, or pale brown _________________________ R. miliacea<br />

29. Inflorescence branches ascending, <strong>the</strong> inflorescence not as above; achenes brown <strong>to</strong> black.<br />

30. Spikelets 4–6(–7.5) mm long; achenes 2–2.5 mm long; <strong>plants</strong> <strong>of</strong> droughty sandhills,<br />

typically with longleaf pines ____________________________________________________ R. grayi<br />

30. Spikelets usually 2–4 mm long (<strong>to</strong> 5 mm long in R. caduca); achenes usually 1–1.8 mm<br />

long; <strong>plants</strong> <strong>of</strong> wet <strong>to</strong> seasonally moist habitats.<br />

31. Perianth bristles shorter than <strong>the</strong> achenes.<br />

32. Surface <strong>of</strong> achenes with horizontal rows <strong>of</strong> cells nearly circular in outline, brown<br />

<strong>to</strong> dark brown or almost black; summit <strong>of</strong> achenes <strong>of</strong>ten abruptly narrowed,<br />

with a distinct, smooth, collar-like ring or ridge fitted against <strong>the</strong> tubercle _______ R. harveyi<br />

32. Surface <strong>of</strong> achenes with horizontal rows <strong>of</strong> vertically oriented cells rectangular<br />

in outline, light brown <strong>to</strong> brown; summit <strong>of</strong> achenes usually gradually narrowed<br />

<strong>to</strong> tubercle base, usually lacking a collar-like ring or ridge.<br />

33. Achenes 1–1.2 mm long, 0.8–1 mm wide; species possibly present in East<br />

TX __________________________________________________________ R. microcarpa<br />

33. Achenes 1.2–1.8 mm long, 1–1.5 mm wide; species widespread and abundant<br />

in East TX.

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