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832 POACEAE/ARTHRAXON A. purpurea; however, “the length of its glumes, the width of its lemma apex, and the length and thickness of its awns distinguish it from all the other varieties” (Allred 2003d). Allred (2003d) noted that the callus and awns are particularly problematic for sheep and cattle. var. nealleyi (Vasey) Allred, (for Greenleaf Alley Nealley, 1864–1896, botanist), BLUE THREEAWN, NEALLEY’S THREEAWN. Panicles contracted; glumes unequal, the lower glume about half as long as upper; lemma 7–12(–13) mm long, usually slightly (1–3 mm) longer than upper glume, narrowing to ca. 0.1 mm wide, the lemma apex twisted into awn column 1–4 mm long; lemma awns slightly unequal, 15–20(–30) mm long. Grasslands on rocky, usually limestone soils; Travis Co. (BRIT; Carr 2002a) near w margin of East TX; w 2/3 of TX; mainly sw U.S. from KS and TX w to CA, also IL and VT. May–Oct. [A. glauca (Nees) Walp.] Turner et al. (2003) combined this taxon and A. purpurea var. wrightii and treated them as A. glauca. However, we are following the recent treatment in Flora of North America (Allred 2003d) in recognizing them as varieties of A. purpurea. According to Allred (2003d), “Although var. nealleyi is more distinct than the other varieties, having tight tufts of foliage exceeded by narrow, straw-colored panicles, it grades into var. purpurea, and the panicles resemble those of var. wrightii.” var. purpurea, (purple), PURPLE THREEAWN, PURPLE NEEDLE GRASS. Panicles open, curving, the branches flexuous; glumes unequal, the second 11–15 mm long, to twice the length of the first; lemma usually 10–12 mm long, narrowing to 0.1–0.3 mm wide at apex; lemma awns about equal, 3–4.5 cm long. Sandy or rocky soils; Post Oak Savannah s and w through much of w 3/4 of TX; sw U.S. from AR and LA w to CA. Apr–Oct. var. wrightii (Nash) Allred, (for Charles Wright, 1811–1885, TX plant collector), WRIGHT’S THREEAWN, WRIGHT’S TRIPLEAWN GRASS. Panicles contracted; glumes unequal, the second 11–15 mm long, the first 0.5–0.75 as long as second; lemma equaling or exceeding upper glume, usually 11–15.5 mm long, narrowing to 0.2–0.3 mm wide at apex; lemma awns about equal, 15–30 mm long, the central occasionally longer than laterals. Calcareous soils; Blackland Prairie s and w through much of w 3/4 of TX; sw U.S. from OK and TX w to CA. May–Oct. [A. wrightii Nash] According to Allred (2003d), “It is the most robust variety of A. purpurea, and has dark, stout awns and long panicles. It may be confused with var. nealleyi, which has narrower lemmas and awns and a light-colored panicle, but it also intergrades with var. purpurea …” Aristida ramosissima Engelm. ex A. Gray, (much-branched), S-CURVE THREEAWN. Annual, much-branched, similar to A. oligantha (inflorescence few-flowored; spikelets large); lower glume 11–20 mm long, upper glume usually 16–25 mm long, including 3–7 mm long awn; lemma (8–)14–25 mm long; central lemma awn 9–28 mm long, with a semicircular bend at base; lateral lemma awns very short, 0–7 mm long, erect or nearly so. Sandy soils; roadsides, fields, open dry areas; Robertson, Van Zandt (BRIT), and Freestone (Turner et al. 2003) cos. in the Post Oak Savannah; rare in TX (e.g., Gould 1975 indicated that he had seen no TX collections); also reported for the Pineywoods and Edwards Plateau by Hatch (2002); c U.S. from IN to IA s to LA and TX. Late summer–fall. While not officially designated as such (e.g., TOES 1993; Carr 2002d; Poole et al. 2002), given its limited distribution in the state, we consider this species to be of conservation concern in TX. � ARTHRAXON P. Beauv. CARP GRASS AA genus of 7 species (Thieret 2003c) of small grasses native to tropical and subtropical areas of the Old World. Like all members of the Andropogoneae, it is characterized by C4 photosyn thesis (Watson & Dallwitz 1999; Kellogg 2000a). While monophyletic, delimitation of species within the genus is “quite difficult” (van Welzen 1993). “The spikelets break off easily and remain stuck to clothes and hair…” (van Welzen 1993). (Greek: arthron, joint, and axon, axis, in reference to the jointed inflorescence axis) (subfamily Panicoideae, tribe Andropogoneae) REFERENCES: Kiger 1971; van Welzen 1981, 1993; Brown & Schultz 1991; Thieret 2003c.
ILLUSTRATED FLORA OF EAST TEXAS 833 Andropogon virginicus Anthenantia texana Anthenantia villosa Anthoxanthum odoratum Aristida basiramea Aristida desmantha Aristida dichotoma Aristida lanosa Aristida longespica (both vars.) Aristida oligantha Aristida palustris Aristida purpurascens (both vars.)
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KEYS TO THE VASCULAR PLANTS 309 KEY
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KEY TO FERNS AND SIMILAR PLANTS 311
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Key to Families of Gymnosperms KEY
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20. Flowers in a single, small (4-4
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38. Stamens 6 per flower. 39. Peria
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57. Leaves reduced to scales only;
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FERNS AND SIMILAR PLANTS (PTERIDOPH
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ISOETES/ISOETACEAE 323 DIVISION LYC
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Psilotum nudum [KNO] Isoetes butler
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LYCOPODIELLA/LYCOPODIACEAE 327 Beit
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SELAGINELLA/SELAGINELLACEAE 329 Pse
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ILLUSTRATED FLORA OF EAST TEXAS 331
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Palhinhaea cernua [CLU] Selaginella
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leaf blade petiole rhizome root sca
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ASPLENIUM L. SPLEENWORT ASPLENIUM/A
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WOODWARDIA/BLECHNACEAE 339 1. Plant
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MEGASPORE Woodwardia areolata [LUN]
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DRYOPTERIDACEAE 343 AThe family as
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Anemia mexicana Azolla caroliniana
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DRYOPTERIS/DRYOPTERIDACEAE 347 Carr
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WOODSIA/DRYOPTERIDACEAE 351 Tectari
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MARSILEA/MARSILEACEAE 353 etation (
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BOTRYCHIUM/OPHIOGLOSSACEAE 357 REFE
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Nephrolepis exaltata Onoclea sensib
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OPHIOGLOSSUM/OPHIOGLOSSACEAE 361 Op
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PTERIDACEAE 365 and pous or podium,
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Botrychium virginianum Ophioglossum
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CERATOPTERIS/PTERIDACEAE 371 Astrol
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Argyrochosma dealbata [LUN] Astrole
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Cheilanthes feei [LUN] Cheilanthes
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Astrolepis integerrima Astrolepis s
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SALVINIACEAE 379 fern, derived from
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SALVINIA/SALVINIACEAE 381 NM, and N
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Pellaea ovata Pellaea wrightiana IL
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Thelypteris dentata [SM3] STERILE F
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GYMNOSPERMS EPHEDRACEAE 389 AThe te
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DIVISION PINOPHYTA CONIFERS CUPRESS
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JUNIPERUS/CUPRESSACEAE 393 allergen
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Ephedra antisyphilitica [LUN] Junip
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PINUS/PINACEAE 397 clusters at the
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� � � � � � Pinus echin
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Ephedra antisyphilitica Juniperus a
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MAGNOLIOPHYTA 403 consumed in large
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CLASS MONOCOTYLEDONAE MONOCOTYLEDON
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AGAVACEAE 407 produced in North Ame
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HESPERALOE/AGAVACEAE 409 for their
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Acorus calamus [GLE] Hesperaloe eng
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YUCCA/AGAVACEAE 413 snakebite (Verh
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7. Leaves straight or nearly so, la
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Manfreda virginica subsp. lata [HEA
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Manfreda maculosa Manfreda sileri I
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ALISMA/ALISMATACEAE 423 1. Pistils
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Yucca rupicola [LYN] Yucca treculea
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SAGITTARIA/ALISMATACEAE 427 long te
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Echinodorus tenellus [FLN, RHO] Sag
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Yucca treculeana ILLUSTRATED FLORA
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ALLIUM/ALLIACEAE 433 pounds are ant
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ALLIUM/ALLIACEAE 435 Allium ampelop
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STAMENS ILLUSTRATED FLORA OF EAST T
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CAPSULE Allium canadense var. mobil
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var. fraseri-▲ var. hyacinthoides
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AMARYLLIDACEAE 443 and edible (Baye
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COOPERIA/AMARYLLIDACEAE 445 REFEREN
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BULBILS CAPSULE Allium sativum [ANO
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HABRANTHUS/AMARYLLIDACEAE 449 obser
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CAPSULES SEED Habranthus tubispathu
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LYCORIS/AMARYLLIDACEAE 455 usually
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Hymenocallis liriosme [HEA] Leucoju
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Narcissus jonquilla [NIC] Narcissus
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ANTHERICACEAE 461 2000), Anderson,
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ARACEAE 463 hemiepiphytes, and aqua
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ARISAEMA/ARACEAE 465 has been disag
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ORONTIUM/ARACEAE 469 down the insid
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Echeandia flavescens [MCV] Colocasi
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ARECACEAE 473 oped rhizomes present
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ASPARAGACEAE 475 spreading from cul
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BROMELIACEAE 477 jointed pedicels;
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Xanthosoma sagittifolium [BA1] Saba
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APTERIA/BURMANNIACEAE 481 FAMILY RE
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S. minor � S. mexicana-� ILLUST
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UVULARIA/COLCHICACEAE 485 blurred,
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ANTHER CAPSULE PISTIL Apteria aphyl
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CALLISIA/COMMELINACEAE 489 the Haem
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COMMELINA/COMMELINACEAE 491 4. Spat
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SINGLE FLOWER INFLORESCENCE Callisi
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TRADESCANTIA/COMMELINACEAE 495 & Tv
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FRUITING INFLORESCENCE Murdannia nu
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Tradescantia paludosa [AND] Tradesc
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CYPERACEAE 507 become split by grow
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CYPERACEAE 509 10. Scales of spikel
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BULBOSTYLIS/CYPERACEAE 511 1. Spike
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CAREX/CYPERACEAE 517 The following
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CAREX/CYPERACEAE 519 of leaf sheath
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CAREX/CYPERACEAE 521 37. Perigynia
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CAREX/CYPERACEAE 523 58. Achene sho
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CAREX/CYPERACEAE 525 85. Larger per
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Carex abscondita [MAC] Carex albica
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CAREX/CYPERACEAE 531 persists in li
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Carex decomposita [MAC] Carex digit
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Carex triangularis [MAC] Carex tetr
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CYPERUS/CYPERACEAE 559 1974; Baijan
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CYPERUS/CYPERACEAE 561 21. Leaf bla
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CYPERUS/CYPERACEAE 563 fresh; achen
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CYPERUS/CYPERACEAE 565 tively more
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CYPERUS/CYPERACEAE 573 Cyperus invo
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Cyperus esculentus [REE] Cyperus fi
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CYPERUS/CYPERACEAE 577 around the s
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Cyperus iria [GWO] Cyperus lanceola
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Cyperus surinamensis [BT2] Cyperus
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ELEOCHARIS/CYPERACEAE 587 5. Culms
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ELEOCHARIS/CYPERACEAE 589 27. Spike
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ELEOCHARIS/CYPERACEAE 591 43. Scale
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ELEOCHARIS/CYPERACEAE 593 compressa
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ELEOCHARIS/CYPERACEAE 601 ing mats;
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FIMBRISTYLIS Vahl FIMBRY FIMBRISTYL
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Fimbristylis annua [SID] Fimbristyl
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REFERENCES: Coville 1890; Bush 1905
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RHYNCHOSPORA/CYPERACEAE 621 Lipocar
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RHYNCHOSPORA/CYPERACEAE 623 10. Per
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RHYNCHOSPORA/CYPERACEAE 625 34. Cul
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Lipocarpha aristulata [GWO] Lipocar
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SCHOENOPLECTUS/CYPERACEAE 641 exten
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Schoenoplectus pungens [MAS] Schoen
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SCLERIA/CYPERACEAE 649 5. Lowest la
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SCLERIA/CYPERACEAE 651 with a pair
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ERIOCAULON/ERIOCAULACEAE 657 1. Pet
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ERIOCAULON/ERIOCAULACEAE 659 bracts
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� Dioscorea bulbifera [HEB] ILLUS
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HYACINTHACEAE 663 orange zone towar
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Scleria pauciflora ILLUSTRATED FLOR
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� � ILLUSTRATED FLORA OF EAST T
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SCHOENOLIRION/HYACINTHACEAE 669 som
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HYDROCHARITACEAE 671 AA small (116
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HYDRILLIA/HYDROCHARITACEAE 673 marg
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OTTELIA Pers. DUCK-LETTUCE OTTELIA/
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� Vallisneria americana [LAM] Ott
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IRIDACEAE 683 inflorescences with 1
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BELAMCANDA Adans. LEOPARD-LILY, BLA
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IRIS/IRIDACEAE 691 6. Cauline leaf
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Iris brevicaulis [YAT] ILLUSTRATED
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SISYRINCHIUM/IRIDACEAE 697 3. Inflo
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Sisyrinchium ensigerum [CUR] Sisyri
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JUNCUS/JUNCACEAE 703 mainly in wet
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JUNCUS/JUNCACEAE 705 yellowish, lea
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Juncus acuminatus [MAS] Juncus anth
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Juncus dichotomus [GWO] Juncus effu
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Juncus polycephalus [GWO] Juncus re
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var. validus-● var. facinatus-�
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LEMNA/LEMNACEAE 721 Pineywoods and
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SPIRODELA/LEMNACEAE 723 Lemna turio
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NOTE 3 VEINS � TOP VIEW Lemna min
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LILIACEAE 729 The lily family (even
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ERYTHRONIUM/LILIACEAE 731 17. Tepal
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Wolffia columbiana [GWO, MIB] Wolff
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LIMNOCHARITACEAE 735 1. Perianth wh
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THALIA/MARANTACEAE 737 pollen prese
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MELANTHIACEAE 739 AA small family (
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Lilium philippinense [GCH] SEED FOL
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VERATRUM/MELANTHIACEAE 745 circumsc
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Zigadenus glaberrimus [GLE] Schoeno
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ALETRIS/NARTHECIACEAE 749 Tofieldia
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DASYLIRION Zucc. SOTOL, SHAGGY-LILY
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FLOWER Aletris aurea [CO1] Aletris
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ORCHIDACEAE 755 45 occur in East TX
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CALOPOGON/ORCHIDACEAE 757 5-25 flow
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CLEISTES/ORCHIDACEAE 759 long; lip
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CYPRIPEDIUM/ORCHIDACEAE 761 while t
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Stenanthium gramineum Toxicoscordio
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Calopogon tuberosus [FNA] Corallorh
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HEXALECTRIS/ORCHIDACEAE 767 rain. A
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Habenaria quinqueseta [LUN] Habenar
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PLATANTHERA/ORCHIDACEAE 771 PLATANT
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PLATANTHERA/ORCHIDACEAE 773 Platant
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SPIRANTHES/ORCHIDACEAE 779 mm wide,
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Page 477 and 478: ILLUSTRATED FLORA OF EAST TEXAS 785
Page 481 and 482: SPIRANTHES/ORCHIDACEAE 789 narrowes
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Page 487 and 488: LEAF collar rhizome blade ligule sh
Page 489 and 490: POACEAE 797 Dissection is usually n
Page 491 and 492: POACEAE 799 rower and often differe
Page 493 and 494: POACEAE 801 6. Lemmas awned, the tw
Page 495 and 496: POACEAE 803 30. Lemma veins none to
Page 497 and 498: POACEAE 805 13. Inflorescences narr
Page 499 and 500: POACEAE 807 12. Lemma awnless; infl
Page 501 and 502: POACEAE 809 inflorescence solitary,
Page 503 and 504: POACEAE 811 29. Spikelets loosely a
Page 505 and 506: POACEAE 813 20. Basal leaves usuall
Page 507 and 508: AEGILOPS/POACEAE 815 23. Spikelets
Page 509 and 510: AGROSTIS/POACEAE 817 Agrostis hyema
Page 513 and 514: ANDROPOGON/POACEAE 821 colonizing e
Page 517 and 518: ANTHENANTIA/POACEAE 825 to be the e
Page 521 and 522: ARISTIDA/POACEAE 829 9. Lemma thick
Page 523: ARISTIDA/POACEAE 831 BRIT), Newton
Page 527 and 528: Anthoxanthum aristatum [BB2] Anthox
Page 531 and 532: AXONOPUS/POACEAE 839 duced to the U
Page 533 and 534: Avena fatua [HI1] Arundo donax [LYN
Page 535 and 536: BOTHRIOCHLOA/POACEAE 843 var. barbi
Page 537 and 538: Aristida purpurea (all vars.) ILLUS
Page 539 and 540: Axonopus compressus [HI1] Axonopus
Page 541 and 542: BOUTELOUA/POACEAE 849 state grass b
Page 543 and 544: Bothriochloa ischaemum var. songari
Page 547 and 548: Bouteloua pectinata [BOT, HEA] Bout
Page 549 and 550: BROMUS/POACEAE 857 8. Lemmas and gl
Page 553 and 554: BUCHLOE/POACEAE 861 ticularly in th
Page 557 and 558: CHASMANTHIUM/POACEAE 865 [C. caroli
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Page 565 and 566: Chasmanthium sessiliflorum [USB] Ch
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Cynodon dactylon [REE] Cynosurus ec
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DICHANTHELIUM/POACEAE 885 AA genus
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DICHANTHELIUM/POACEAE 887 20. Inter
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DICHANTHELIUM/POACEAE 889 26. Leaf
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Danthonia spicata [USB] Desmazeria
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DICHANTHELIUM/POACEAE 897 and Harri
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Dichanthelium acuminatum (all vars.
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DICHANTHELIUM/POACEAE 905 that this
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DIGITARIA/POACEAE 909 2. Main axis
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Dichanthelium sphaerocarpon [HI] IL
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DIGITARIA/POACEAE 913 with papilla-
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Digitaria bicornis [HEB] ILLUSTRATE
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Digitaria insularis [HI1] Digitaria
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EHRHARTA/POACEAE 921 spikelets (2.5
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ELIONURUS/POACEAE 923 Since this co
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Echinochloa crus-gali [REE] Echinoc
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ENTEROPOGON Nees UMBRELLA GRASS ENT
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ERAGROSTIS/POACEAE 931 8. Infloresc
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ERAGROSTIS/POACEAE 933 12. Pedicels
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Elymus repens [HI1] Elymus smithii
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Eragrostis japonica [USB] ILLUSTRAT
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ERIONEURON/POACEAE 945 Shaw and Sme
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� � � ILLUSTRATED FLORA OF EA
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FESTUCA/POACEAE 951 rachilla glabro
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GYMNOPOGON/POACEAE 955 Glyceria sep
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HILARIA/POACEAE 957 REFERENCES: Eme
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ILLUSTRATED FLORA OF EAST TEXAS Fes
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KOELERIA/POACEAE 963 Andropogoneae,
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Heteropogon melanocarpus [HI1] Hord
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LEPTOCHLOA/POACEAE 967 Leersia oryz
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ILLUSTRATED FLORA OF EAST TEXAS Koe
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ILLUSTRATED FLORA OF EAST TEXAS Hor
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LOLIUM/POACEAE 973 Charmet et al. 1
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Leptochloa nealleyi [USB] Lolium ri
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MICROSTEGIUM/POACEAE 977 AA variabl
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ILLUSTRATED FLORA OF EAST TEXAS Lep
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Lolium temulentum [USB] Melica muti
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MUHLENBERGIA/POACEAE 983 10. Lemma
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MUHLENBERGIA/POACEAE 985 and Prairi
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ILLUSTRATED FLORA OF EAST TEXAS Mon
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ILLUSTRATED FLORA OF EAST TEXAS Mic
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 993 fact that gener
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PANICUM/POACEAE 995 17. Spikelets 2
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Oryza sativa [HI1] Panicum anceps [
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 1001 Panicum hallii
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Panicum trichoides [HI2] ILLUSTRATE
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PASPALUM/POACEAE 1009 5. Spikelets
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PASPALUM/POACEAE 1011 27. Axils of
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Panicum virgatum [RCA] Paspalum acu
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PASPALUM/POACEAE 1019 loam soils; w
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PENNISETUM/POACEAE 1025 unit as in
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Paspalum repens [USB] ILLUSTRATED F
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PHALARIS/POACEAE 1029 landscapes (B
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Paspalum pubiflorum (both vars.) IL
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Pennisetum purpureum [HUT] Penniset
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POA/POACEAE 1035 around awn base; l
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POA/POACEAE 1037 long. Forests, flo
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Page 737 and 738:
SACCHARUM/POACEAE 1045 1. Awn of le
Page 739 and 740:
SCHEDONNARDUS/POACEAE 1047 lower gl
Page 741 and 742:
Page 743 and 744:
Page 745 and 746:
SETARIA/POACEAE 1053 Paspalidium (W
Page 747 and 748:
SETARIA/POACEAE 1055 14. Leaf blade
Page 749 and 750:
Schizachyrium tenerum [HI1, USB] Sc
Page 751 and 752:
Page 753 and 754:
Page 755 and 756:
SORGHUM/POACEAE 1063 2000) indicate
Page 757 and 758:
SPARTINA/POACEAE 1065 Americas it [
Page 759 and 760:
Sorghastrum elliottii [HI1] Sorghas
Page 761 and 762:
Page 763 and 764:
SPOROBOLUS/POACEAE 1071 7. Lemma ap
Page 765 and 766:
Page 767 and 768:
STEINCHISMA/POACEAE 1075 Sporobolus
Page 769 and 770:
Sphenopholis obtusata (both vars.)
Page 771 and 772:
Sporobolus neglectus [HI1] Sporobol
Page 773 and 774:
TRICHONEURA/POACEAE 1081 guish it f
Page 775 and 776:
Stenotaphrum secundatum [USB] Trich
Page 777 and 778:
Page 779 and 780:
Page 781 and 782:
TRISETUM/POACEAE 1089 low areas; ne
Page 783 and 784:
UROCHLOA/POACEAE 1091 AA C4 genus o
Page 785 and 786:
Page 787 and 788:
Tridens eragrostoides Tridens flavu
Page 789 and 790:
VULPIA/POACEAE 1097 2. Lemmas (exce
Page 791 and 792:
Page 793 and 794:
Page 795 and 796:
ZIZANIOPSIS/POACEAE 1103 (1999) (po
Page 797 and 798:
Page 799 and 800:
EICHHORNIA/PONTEDERIACEAE 1107 REFE
Page 801 and 802:
Zoysia japonica [HOL] Zoysia pacifi
Page 803 and 804:
POTAMOGETON/POTAMOGETONACEAE 1111 1
Page 805 and 806:
POTAMOGETON/POTAMOGETONACEAE 1113 P
Page 807 and 808:
Page 809 and 810:
Page 811 and 812:
SMILAX/SMILACACEAE 1119 times woody
Page 813 and 814:
SMILAX/SMILACACEAE 1121 30(-60) mm
Page 815 and 816:
Stuckenia pectinata [MAS] ILLUSTRAT
Page 817 and 818:
Page 819 and 820:
THEMIDACEAE 1127 cover for various
Page 821 and 822:
Smilax rotundifolia [JEM] ILLUSTRAT
Page 823 and 824:
TRILLIUM/TRILLIACEAE 1131 Schilling
Page 825 and 826:
TRILLIUM/TRILLIACEAE 1133 taxonomic
Page 827 and 828:
Triantha racemosa [GLE] Trillium lu
Page 829 and 830:
TYPHA/TYPHACEAE 1137 REFERENCES: Wi
Page 831 and 832:
XYRIS/XYRIDACEAE 1139 cone-like spi
Page 833 and 834:
XYRIS/XYRIDACEAE 1141 11. Plant bas
Page 835 and 836:
Sparganium americanum Typha latifol
Page 837 and 838:
Typha domingensis[ BT3] Typha latif
Page 839 and 840:
Xyris drummondii [SID] Xyris fimbri
Page 841 and 842:
Xyris smalliana [SID] ILLUSTRATED F
Page 843:
Xyris caroliniana Xyris difformis (
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keys to the vascular plants of east texas - Botanical Research ...

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