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STEINCHISMA/POACEAE<br />

1075<br />

Sporobolus pyramidatus (Lam.) Hitchc., (pyramidal), WHORLED DROPSEED, TARGET DROPSEED.<br />

Annual or short-lived perennial; culms 10–35(–60) cm tall; panicles 3–15(–18) cm long, becoming<br />

pyramidal and exserted at maturity, <strong>the</strong> lower branches in whorls <strong>of</strong> usually 7–12(–rarely<br />

more); spikelets 1.5–2 mm long. Open, disturbed sites; Navarro (BRIT), Anderson, Bastrop, Bexar,<br />

Brazos, Hunt, and Tyler (Turner et al. 2003) cos.; widely scattered in TX, but more common in<br />

<strong>the</strong> s part <strong>of</strong> <strong>the</strong> state; c U.S. from MO s <strong>to</strong> LA w <strong>to</strong> UT and AZ, also FL. Mar–Nov. [Agrostis<br />

pyramidata Lam., S. patens Swallen, S. pulvinatus Swallen] While this taxon is sometimes included<br />

in <strong>the</strong> Old World S. coromandelianus (Retz.) Kunth (e.g., Kartesz 1999; Hatch 2002), we<br />

are following Peterson et al. (2003) in treating S. pyramidatus as a separate species. They noted,<br />

however, that <strong>the</strong> two are morphologically “very similar … suggesting that <strong>the</strong>y are related.”<br />

Sporobolus silveanus Swallen, (for William Arents Silveus, 1875–1953, TX botanist, at<strong>to</strong>rney,<br />

and author <strong>of</strong> Texas Grasses), SILVEUS’ DROPSEED. Perennial; culms 0.7–1.2 m tall; panicles somewhat<br />

open, 20–50 cm long, (5–)10–12(–15) cm wide; spikelets purple, (4–)4.5–7(–7.2) mm long.<br />

Sandy soils, prairies, wet <strong>to</strong> mesic pine woodlands, forest openings; scattered in East TX in Hardin,<br />

Jasper, Rains (BRIT), Angelina, Brazos, Harris, Lamar (Tridens Prairie), Orange, Tyler, and<br />

Van Zandt (Weakley & Peterson 1998) cos.; also n Gulf Prairies and Marshes; w LA, se OK, and<br />

East TX (Weakley & Peterson 1998); Peterson et al. (2003) also mapped a single county in FL<br />

and a single county in GA. Jul–Nov. Brown (1993) concluded that all TX material previously<br />

identified as <strong>the</strong> related species S. heterolepis (A. Gray) A. Gray was actually S. silveanus. We are<br />

following Brown (1993) in not considering S. heterolepis <strong>to</strong> be a member <strong>of</strong> <strong>the</strong> TX flora.<br />

Sporobolus vaginiflorus (Torr. ex A. Gray) A.W. Wood, (with flowers in <strong>the</strong> sheaths), POVERTY<br />

DROPSEED, SOUTHERN POVERTY GRASS, POVERTY GRASS. Annual; culms 15–50(–70) cm tall;<br />

panicles terminal and axillary, contracted, 1–5 cm long, 2–5 mm wide, usually partially enclosed<br />

within subtending sheaths; spikelets usually 3–5(–6) mm long; lemma <strong>of</strong>ten mottled<br />

with dark purple, with short appressed pubescence. Disturbed sites, roadsides, fields, sandy or<br />

clay soils, limes<strong>to</strong>ne outcrops; widespread in e 1/2 <strong>of</strong> TX, scattered fur<strong>the</strong>r w; scattered in s<br />

Canada and throughout <strong>the</strong> e U.S. w <strong>to</strong> ND and TX, also AZ, ID, and NM. Aug–Nov. This species<br />

is similar <strong>to</strong> and apparently related <strong>to</strong> S. ozarkanus; see discussion under that species.<br />

Sporobolus heterolepis (A. Gray) A. Gray, PRAIRIE DROPSEED, has <strong>of</strong>ten been reported for TX (e.g.,<br />

Correll & Johns<strong>to</strong>n 1970; Gould 1975; Hatch et al. 1990). Brown (1993), however, concluded that<br />

all TX material identified as this species was actually S. silveanus. We are following Brown<br />

(1993), Hatch (2002), and Turner et al. (2003) in considering S. heterolepis not <strong>to</strong> be a member<br />

<strong>of</strong> <strong>the</strong> TX flora. However, several old collections are still listed incorrectly as S. heterolepis on<br />

<strong>the</strong> Digital Flora <strong>of</strong> Texas Herbarium Specimen Browser (2002) and were apparently picked up<br />

in <strong>the</strong> recent Flora <strong>of</strong> North America treatment (Peterson et al. 2003).<br />

STEINCHISMA Raf. GAPING PANIC GRASS<br />

AA New World genus <strong>of</strong> 5–6 species <strong>of</strong> tropical <strong>to</strong> warm-temperate regions, growing typically in<br />

moist or wet, usually open, sandy areas (Freckmann & Lelong 2003c). It has traditionally been<br />

included in Panicum (e.g., Correll & Johns<strong>to</strong>n 1970; Hatch et al. 1990), sometimes in <strong>the</strong> subgenus<br />

Steinchisma (Zuloaga 1987). Recent studies (e.g., Zuloaga et al. 2000) support its recognition<br />

as a separate genus (Freckmann & Lelong 2003c). Pho<strong>to</strong>syn<strong>the</strong>sis is intermediate between<br />

that <strong>of</strong> C3 and C4 <strong>plants</strong> (Freckmann & Lelong 2003c). (Greek: steinos, narrow, and chasma,<br />

yawning, presumably in reference <strong>to</strong> <strong>the</strong> gaping glumes and somewhat narrow spikelets when<br />

compared <strong>to</strong> Panicum—Freckmann & Lelong 2003c) (subfamily Panicoideae, tribe Paniceae)<br />

REFERENCES: Zuloaga et al. 1992, 1998. Freckmann & Lelong 2003c.<br />

Steinchisma hians (Elliott) Nash, (gaping), GAPING PANICUM, GAPING GRASS, GAPING PANIC GRASS.<br />

Perennial with culms erect or decumbent at base, <strong>to</strong> 75 cm tall; nodes glabrous or scabrous; leaf<br />

sheaths glabrous or with hairs on upper margins; ligule a short membrane ca. 0.5 mm long; leaf

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