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SCHEDONNARDUS/POACEAE<br />

1047<br />

lower glume ca. 1–1.7 mm long, � 1/3 as long as second, <strong>of</strong>ten 1/4 or less as long; lemma <strong>of</strong> fertile<br />

floret ca. 1.5–2 mm long. Moist sand, pond margins, along streams, and in ditches; widely<br />

scattered in East TX w <strong>to</strong> Lamar, Leon, Wood (BRIT), and Guadalupe (Turner et al. 2003) cos;<br />

also n Gulf Prairies and Marshes; se U.S. from NJ s <strong>to</strong> FL w <strong>to</strong> MO, OK, and TX, also MA and ME.<br />

Late Aug–Nov. [Panicum aquaticum Bosc ex Spreng.]<br />

SCHEDONNARDUS Steud. TUMBLE GRASS, TEXAS CRAB GRASS<br />

AA monotypic C4 genus <strong>of</strong> North America that has recently been introduced in<strong>to</strong> Argentina<br />

(Snow 2003b). Some authorities consider it related <strong>to</strong> Gymnopogon (Clay<strong>to</strong>n & Renvoize 1986),<br />

but recent molecular evidence (Hilu & Alice 2001) questions this conclusion. The inflorescence<br />

acts as a tumbleweed, and Mabberley (1997) noted that this species is a conspicuous feature <strong>of</strong><br />

deserted <strong>to</strong>wns in Western films. (Greek: schedon, near, and Nardus, from its resemblance <strong>to</strong><br />

that grass genus) (subfamily Chloridoideae, tribe Cynodonteae)<br />

REFERENCE: Snow 2003b.<br />

Schedonnardus paniculatus (Nutt.) Trel., (with flowers in panicles), TUMBLE GRASS, TEXAS CRAB<br />

GRASS. Tufted perennial 8–50(–70) cm tall or long; leaves crowded <strong>to</strong>ward base <strong>of</strong> plant; leaf<br />

sheaths compressed, keeled, glabrous; ligule a membrane 1–4 mm long; leaf blades 1–3 mm<br />

wide, spirally twisted on drying; inflorescence as long as or longer than <strong>the</strong> leafy portion <strong>of</strong> <strong>the</strong><br />

culm, spreading <strong>to</strong> partly decumbent, finally breaking away in its entirety and acting as a tumbleweed,<br />

<strong>the</strong> main axis becoming loosely coiled, with a few widely spaced, spicate branches;<br />

spikelets appressed, in 2 rows along one side <strong>of</strong> each branch, not overlapping or only slightly so,<br />

with 1 floret, unawned, 3–5.5 mm long, disarticulating above <strong>the</strong> glumes; glumes unequal, <strong>the</strong><br />

upper longer. Prairies, disturbed areas, various soils; throughout TX; sc Canada and c U.S. from<br />

MN s <strong>to</strong> LA w <strong>to</strong> MT and AZ, also CA. This was a minor member <strong>of</strong> original prairie which increases<br />

under disturbance; ra<strong>the</strong>r common on disturbed or overgrazed sites. Apr–Oct.<br />

SCHIZACHYRIUM Nees BLUESTEM, FALSE BLUESTEM<br />

Ours perennials; ligule membranous, 3 mm or less long, sometimes fringed; flowering culms<br />

much-branched, each leafy branch terminating in a spicate raceme (<strong>the</strong>se sometimes crowded<br />

<strong>to</strong>ge<strong>the</strong>r and sometimes appearing axillary due <strong>to</strong> <strong>the</strong> shortness <strong>of</strong> <strong>the</strong> branches <strong>of</strong> <strong>the</strong> flowering<br />

stem); spikelets in pairs <strong>of</strong> one sessile and one pedicellate; sessile spikelets 2-flowered, <strong>the</strong><br />

upper floret fertile, with <strong>to</strong>o<strong>the</strong>d or cleft lemma, <strong>the</strong> lower floret neuter; pedicellate spikelets<br />

staminate or neuter; disarticulation at base <strong>of</strong> sessile spikelet so that associated pedicel and section<br />

<strong>of</strong> inflorescence axis fall with sessile spikelet.<br />

AA mainly tropical and subtropical genus <strong>of</strong> ca. 60 species (Wipff 2003k), particularly in<br />

savannahs. These <strong>plants</strong> were formerly treated along with Bothriochloa in a more inclusive<br />

Andropogon (e.g., Hitchcock 1951). However, most species <strong>of</strong> Schizachyrium differ from <strong>the</strong>se<br />

o<strong>the</strong>r two genera in having only one raceme per peduncle (Wipff 2003k). In addition,<br />

Schizachyrium species have raceme internodes with cupulate tips and convex lower glumes<br />

veined between <strong>the</strong> keels—Andropogon species lack <strong>the</strong>se features (Wipff 2003k). The common<br />

name, BLUESTEM, is derived from <strong>the</strong> bluish green color <strong>of</strong> <strong>the</strong> glaucous foliage <strong>of</strong> some <strong>plants</strong>.<br />

Like all members <strong>of</strong> <strong>the</strong> Andropogoneae, Schizachyrium is characterized by C4 pho<strong>to</strong>syn<strong>the</strong>sis<br />

(Kellogg 2000a). Several species are important native pasture grasses while some are significant<br />

weeds (Watson & Dallwitz 1992). (Greek: schizo, <strong>to</strong> divide or split, and achna, chaff, referring <strong>to</strong><br />

<strong>the</strong> <strong>to</strong>o<strong>the</strong>d lemma) (subfamily Panicoideae, tribe Andropogoneae)<br />

REFERENCES: Shinners 1954; Gould 1967; Grelen 1974; Türpe 1984; Carmen & Briske 1985; Butler &<br />

Briske 1988; Gandhi 1989; Briske & Anderson 1992; Gandhi & Smeins 1996; Wipff 1996a, 2003k.<br />

1. Pedicels (<strong>of</strong> pedicellate spikelets) ciliate with long silvery hairs at l<strong>east</strong> on <strong>the</strong> upper two-thirds;<br />

raceme internodes relatively thin and flexuous, ciliate with long silvery hairs <strong>the</strong> entire length or

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