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GYMNOPOGON/POACEAE<br />

955<br />

Glyceria septentrionalis Hitchc., (nor<strong>the</strong>rn), FLOATING MANNA GRASS, EASTERN MANNA GRASS,<br />

Plant glabrous; culms ca. 1–1.8 m tall, <strong>of</strong>ten rooting near base; ligule ca. 3–6(–10) mm long; leaf<br />

blades usually 4–10 mm wide; inflorescence usually 18–40(–50) cm long, <strong>the</strong> branches short<br />

and rigid. Wet areas, <strong>of</strong>ten in woods; Bowie, Brazos, Franklin, Trinity, and Robertson (TEX) cos.;<br />

also n Gulf Prairies and Marshes; se Canada and e U.S. w <strong>to</strong> MN and TX. Mar–Jun. [Panicularia<br />

septentrionalis (Hitchc.) E.P. Bicknell]<br />

Glyceria striata (Lam.) Hitchc., (striated, striped), FOWL MANNA GRASS, NERVED MANNA GRASS.<br />

Plant glabrous, <strong>of</strong>ten with short rhizomes; culms 0.4–0.9(–1.5) m tall, usually erect from decumbent<br />

bases; ligule 1–4 mm long; leaf blades usually 2–8 mm wide, rarely wider; inflorescence<br />

10–22(–25) cm long, typically with numerous slender, flexuous branches; spikelets ca. 2–4(–5)<br />

mm long; lemmas usually 1.3–2(–2.5) mm long, glabrous or nearly so, <strong>the</strong> 7 veins usually quite<br />

prominent; an<strong>the</strong>rs 0.3–0.5(–0.9) mm long. Along streams, swamps, marshes, moist forest margins;<br />

Bowie (BAYLU) and Lamar (Carr 1994) cos. in Red River drainage, also Bell (TAES), Travis,<br />

and Washing<strong>to</strong>n (Turner et al. 2003) cos.; rare in e Edwards Plateau and Trans-Pecos; nearly<br />

throughout s Canada and <strong>the</strong> U.S., also disjunct <strong>to</strong> <strong>the</strong> mountains <strong>of</strong> Mexico (Tucker 1996). Apr–<br />

Aug. This species is variable morphologically, but <strong>the</strong> differences appear <strong>to</strong> be “a consequence<br />

<strong>of</strong> environmental, ra<strong>the</strong>r than genetic, fac<strong>to</strong>rs” (Barkworth ined.). It is reported as potentially<br />

cyanogenic (Burlage 1968; Fuller & McClin<strong>to</strong>ck 1986). �<br />

GYMNOPOGON P. Beauv. BEARD GRASS, SKELETON GRASS<br />

Clumped perennials from rhizomes; culms nearly erect <strong>to</strong> decumbent; leaves cauline, distichous<br />

(= 2-ranked), crowded; ligule minute, a fringed membrane or a rim <strong>of</strong> callus tissue, � 1<br />

mm long; leaf blades flat, short, abruptly narrowed and � cordate at base, (resembling those <strong>of</strong><br />

some Panicum species); inflorescence a panicle, mostly ca. as wide as long or wider, with numerous<br />

widely spreading <strong>to</strong> reflexed, spike-like branches (<strong>the</strong>se alternate along <strong>the</strong> main axis<br />

<strong>of</strong> <strong>the</strong> inflorescence), with small beards <strong>of</strong> hairs <strong>of</strong>ten present at base <strong>of</strong> inflorescence branches;<br />

spikelets usually not overlapping, appressed, in 2 rows on 1 side <strong>of</strong> <strong>the</strong> slender, slightly flattened<br />

branches, with 1(–2) perfect lower floret and 1 rudimentary upper floret (usually reduced <strong>to</strong><br />

only a naked or awned pedicel); disarticulation above <strong>the</strong> glumes; glumes subequal (upper<br />

slightly longer), longer than <strong>the</strong> florets; lemma <strong>of</strong> fertile floret with an awn from <strong>the</strong> minutely<br />

notched apex; palea <strong>of</strong>ten slightly longer than lemma.<br />

AA C4 genus <strong>of</strong> ca. 15 species mostly <strong>of</strong> warm areas <strong>of</strong> <strong>the</strong> Americas, with 1 species from India<br />

<strong>to</strong> Thailand (Smith 2003). The foliage is superficially similar <strong>to</strong> that <strong>of</strong> some Panicum species,<br />

while <strong>the</strong> inflorescence is reminiscent <strong>of</strong> Chloris. The genus is related <strong>to</strong> Chloris, but<br />

Gymnopogon can be distinguished by its 2-ranked leaves, alternate inflorescence branches,<br />

glumes longer than <strong>the</strong> florets, and <strong>the</strong> essentially obsolete upper floret (Clay<strong>to</strong>n & Renvoize<br />

1986; Yatskievych 1999). (Greek: gymnus, naked, and pogon, beard, referring <strong>to</strong> <strong>the</strong> reduction <strong>of</strong><br />

<strong>the</strong> abortive flower <strong>to</strong> a bare awn, or <strong>to</strong> <strong>the</strong> naked prolongation <strong>of</strong> <strong>the</strong> rachilla in a number <strong>of</strong><br />

species) (subfamily Chloridoideae, tribe Cynodonteae)<br />

REFERENCES: Smith 1971, 2003.<br />

1. Branches <strong>of</strong> inflorescence bearing scattered spikelets usually <strong>to</strong> <strong>the</strong> base (<strong>the</strong> very base with a<br />

spikelet); rudimentary floret with a distinct awn or sometimes reduced <strong>to</strong> only an awned pedicel;<br />

rhizomes short, knotty or plant merely with a knotty base; lemma <strong>of</strong> fertile floret with awn 4–8<br />

(–12) mm long __________________________________________________________________ G. ambiguus<br />

1. Branches <strong>of</strong> inflorescence without spikelets in <strong>the</strong> basal 1/3 or more <strong>of</strong> <strong>the</strong>ir length (<strong>the</strong> very<br />

base without a spikelet); rudimentary floret awnless, reduced <strong>to</strong> only a naked pedicel; rhizomes<br />

elongate; lemma <strong>of</strong> fertile floret with awn 1–3(–4.5) mm long ____________________________ G. brevifolius

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