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1072 POACEAE/SPOROBOLUS Sporobolus compositus (Poir.) Merr., (compound), HEAD-LIKE DROPSEED, ROUGH DROPSEED, FLAG GRASS. Perennial; culms (20–)60–120(–150) cm tall; panicle 5–30 cm long, at least partly included within leaf sheaths (cleistogamous spikelets often present in axillary panicles); spikelets 4–7.5(–10) mm long. [S. asper (Michx.) Kunth] This species long went under the name S. asper, but because of nomenclatural considerations, S. compositus is the appropriate binomial (Kartesz & Gandhi 1995). The map in Turner et al. (2003) and the one presented here do not separate varieties. According to Peterson et al. (2003), “The Sporobolus compositus complex is a difficult assemblage of forms, perhaps affected by their primarily autogamous breeding system.” Sporobolus clandestinus is sometimes recognized as a variety of this species (e.g., Wipff & Jones 1995); see that species for discussion. However, we are following some recent authors (e.g., Peterson et al. 2003) in recognizing only three varieties in S. compositus. 1. Plants with evident short, creeping, scaly rhizomes _______________________________________ var. macer 1. Plants without creeping, scaly rhizomes. 2. Terminal leaf sheaths 0.8–2(–2.5) mm wide when folded naturally (do not spread out to measure); culms slender, 1–2(–2.5) mm wide near base; primary inflorescence branches 8–18, not crowded _________________________________________________________________ var. drummondii 2. Terminal leaf sheaths (1.3–)1.5–6 mm wide when folded; culms stout, (1.4–)2–5 mm wide near base; primary inflorescence branches 12–35, crowded _________________________ var. compositus var. compositus, TALL DROPSEED, LONG-LEAF RUSH GRASS, ROUGH RUSH GRASS. Cespitose, without rhizomes. Grasslands, disturbed sites; widespread in e 2/3 of TX; s Canada and widespread in the U.S., particularly the e 2/3. Sep–Nov. var. drummondii (Trin.) Kartesz & Gandhi, (for its discoverer, Thomas Drummond, 1780–1835, Scottish botanist and collector in North America), MEADOW DROPSEED. Cespitose, without rhizomes. Grasslands, disturbed sites; widespread in e 1/2 of TX; sc U.S. from KY and GA w to IA and TX. Aug–Nov. [S. asper (Michx.) Kunth var. drummondii (Trin.) Vasey, S. drummondii (Trin.) Vasey] Jones et al. (1997) included this variety in var. compositus. var. macer (Trin.) Kartesz & Gandhi, (thin, meager), MISSISSIPPI DROPSEED. Similar to var. drummondii, but rhizomatous; culms slender, 2(–2.5) mm or less wide; terminal leaf sheath 2.5 mm or less wide when folded. Open woods, margins of woods; Gregg, Titus (BRIT), Anderson, Bastrop, Brazos, Burleson, Camp, Jasper, and Robertson (TAES—annotated by S. Hatch) cos.; sc U.S. from MO and MS w to KS and TX. Aug–Nov. [S. asper (Michx.) Kunth var. macer (Trin.) Shinners, S. macer (Trin.) Hitchc.] Sporobolus cryptandrus (Torr.) A. Gray, (with hidden flowers), SAND DROPSEED, COVERED-SPIKE DROPSEED. Perennial; culms 25–100(–120) cm tall; summit of sheaths with tufts of long white hairs 2–4 mm long; panicles 15–30(–40) cm long, 2–8(–15) cm wide, usually partially enclosed by the subtending sheaths; spikelets 1.5–2.3(–2.8) mm long; lemma ca. as long as or slightly longer than the upper glume. Grasslands, disturbed sites, sandy soils; throughout TX except Pineywoods; s Canada and throughout most of the U.S. May–Nov. The grains were consumed by Native Americans (Mabberley 1987); they were apparently ground into a flour and then baked (Yatskievych 1999). Sporobolus indicus (L.) R. Br., (of India), SMUT GRASS. Perennial; culms 30–100(–120) cm tall; panicles 10–30(–50) cm long; spikelets 1.4–2(–2.6) mm long; pericarp mucilaginous, the seed often sticking instead of falling readily. Moist soils, often in disturbed areas; widespread in e 1/3 of TX, sparsely scattered further w; e U.S. from NY s to FL w to MO, OK, and TX, also CA, MI, and OR. Mar–Dec. Native to Asia. The common name, SMUT GRASS, refers to the infection and blackening of the inflorescences and upper leaves by a fungus (Bipolaris spp.) (Peterson et al. 2003). I
ILLUSTRATED FLORA OF EAST TEXAS 1073 Sphenopholis obtusata var. major [CRO, HI1] Sphenopholis obtusata var. obtusata [GO1] Sporobolus airoides [LEI] Sporobolus clandestinus [HI1] Sporobolus compositus var. compositus [USB] Sporobolus compositus var. drummondii [BB2] Sporobolus compositus var. macer [HI1] Sporobolus cryptandrus [USB] Sporobolus indicus [REE]
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KEYS TO THE VASCULAR PLANTS 309 KEY
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KEY TO FERNS AND SIMILAR PLANTS 311
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Key to Families of Gymnosperms KEY
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20. Flowers in a single, small (4-4
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38. Stamens 6 per flower. 39. Peria
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57. Leaves reduced to scales only;
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FERNS AND SIMILAR PLANTS (PTERIDOPH
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ISOETES/ISOETACEAE 323 DIVISION LYC
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Psilotum nudum [KNO] Isoetes butler
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LYCOPODIELLA/LYCOPODIACEAE 327 Beit
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SELAGINELLA/SELAGINELLACEAE 329 Pse
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ILLUSTRATED FLORA OF EAST TEXAS 331
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Palhinhaea cernua [CLU] Selaginella
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leaf blade petiole rhizome root sca
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ASPLENIUM L. SPLEENWORT ASPLENIUM/A
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WOODWARDIA/BLECHNACEAE 339 1. Plant
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MEGASPORE Woodwardia areolata [LUN]
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DRYOPTERIDACEAE 343 AThe family as
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Anemia mexicana Azolla caroliniana
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DRYOPTERIS/DRYOPTERIDACEAE 347 Carr
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WOODSIA/DRYOPTERIDACEAE 351 Tectari
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MARSILEA/MARSILEACEAE 353 etation (
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BOTRYCHIUM/OPHIOGLOSSACEAE 357 REFE
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Nephrolepis exaltata Onoclea sensib
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OPHIOGLOSSUM/OPHIOGLOSSACEAE 361 Op
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PTERIDACEAE 365 and pous or podium,
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Botrychium virginianum Ophioglossum
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CERATOPTERIS/PTERIDACEAE 371 Astrol
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Argyrochosma dealbata [LUN] Astrole
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Cheilanthes feei [LUN] Cheilanthes
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Astrolepis integerrima Astrolepis s
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SALVINIACEAE 379 fern, derived from
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SALVINIA/SALVINIACEAE 381 NM, and N
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Pellaea ovata Pellaea wrightiana IL
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Thelypteris dentata [SM3] STERILE F
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GYMNOSPERMS EPHEDRACEAE 389 AThe te
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DIVISION PINOPHYTA CONIFERS CUPRESS
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JUNIPERUS/CUPRESSACEAE 393 allergen
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Ephedra antisyphilitica [LUN] Junip
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PINUS/PINACEAE 397 clusters at the
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� � � � � � Pinus echin
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Ephedra antisyphilitica Juniperus a
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MAGNOLIOPHYTA 403 consumed in large
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CLASS MONOCOTYLEDONAE MONOCOTYLEDON
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AGAVACEAE 407 produced in North Ame
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HESPERALOE/AGAVACEAE 409 for their
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Acorus calamus [GLE] Hesperaloe eng
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YUCCA/AGAVACEAE 413 snakebite (Verh
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7. Leaves straight or nearly so, la
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Manfreda virginica subsp. lata [HEA
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Manfreda maculosa Manfreda sileri I
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ALISMA/ALISMATACEAE 423 1. Pistils
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Yucca rupicola [LYN] Yucca treculea
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SAGITTARIA/ALISMATACEAE 427 long te
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Echinodorus tenellus [FLN, RHO] Sag
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Yucca treculeana ILLUSTRATED FLORA
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ALLIUM/ALLIACEAE 433 pounds are ant
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ALLIUM/ALLIACEAE 435 Allium ampelop
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STAMENS ILLUSTRATED FLORA OF EAST T
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CAPSULE Allium canadense var. mobil
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var. fraseri-▲ var. hyacinthoides
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AMARYLLIDACEAE 443 and edible (Baye
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COOPERIA/AMARYLLIDACEAE 445 REFEREN
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BULBILS CAPSULE Allium sativum [ANO
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HABRANTHUS/AMARYLLIDACEAE 449 obser
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CAPSULES SEED Habranthus tubispathu
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LYCORIS/AMARYLLIDACEAE 455 usually
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Hymenocallis liriosme [HEA] Leucoju
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Narcissus jonquilla [NIC] Narcissus
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ANTHERICACEAE 461 2000), Anderson,
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ARACEAE 463 hemiepiphytes, and aqua
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ARISAEMA/ARACEAE 465 has been disag
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ORONTIUM/ARACEAE 469 down the insid
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Echeandia flavescens [MCV] Colocasi
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ARECACEAE 473 oped rhizomes present
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ASPARAGACEAE 475 spreading from cul
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BROMELIACEAE 477 jointed pedicels;
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Xanthosoma sagittifolium [BA1] Saba
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APTERIA/BURMANNIACEAE 481 FAMILY RE
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S. minor � S. mexicana-� ILLUST
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UVULARIA/COLCHICACEAE 485 blurred,
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ANTHER CAPSULE PISTIL Apteria aphyl
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CALLISIA/COMMELINACEAE 489 the Haem
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COMMELINA/COMMELINACEAE 491 4. Spat
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SINGLE FLOWER INFLORESCENCE Callisi
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TRADESCANTIA/COMMELINACEAE 495 & Tv
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FRUITING INFLORESCENCE Murdannia nu
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Tradescantia paludosa [AND] Tradesc
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CYPERACEAE 507 become split by grow
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CYPERACEAE 509 10. Scales of spikel
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BULBOSTYLIS/CYPERACEAE 511 1. Spike
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CAREX/CYPERACEAE 517 The following
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CAREX/CYPERACEAE 519 of leaf sheath
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CAREX/CYPERACEAE 521 37. Perigynia
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CAREX/CYPERACEAE 523 58. Achene sho
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CAREX/CYPERACEAE 525 85. Larger per
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Carex abscondita [MAC] Carex albica
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CAREX/CYPERACEAE 531 persists in li
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Carex decomposita [MAC] Carex digit
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Carex triangularis [MAC] Carex tetr
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CYPERUS/CYPERACEAE 559 1974; Baijan
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CYPERUS/CYPERACEAE 561 21. Leaf bla
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CYPERUS/CYPERACEAE 563 fresh; achen
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CYPERUS/CYPERACEAE 565 tively more
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CYPERUS/CYPERACEAE 573 Cyperus invo
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Cyperus esculentus [REE] Cyperus fi
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CYPERUS/CYPERACEAE 577 around the s
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Cyperus iria [GWO] Cyperus lanceola
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Cyperus surinamensis [BT2] Cyperus
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ELEOCHARIS/CYPERACEAE 587 5. Culms
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ELEOCHARIS/CYPERACEAE 589 27. Spike
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ELEOCHARIS/CYPERACEAE 591 43. Scale
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ELEOCHARIS/CYPERACEAE 593 compressa
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ELEOCHARIS/CYPERACEAE 601 ing mats;
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FIMBRISTYLIS Vahl FIMBRY FIMBRISTYL
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Fimbristylis annua [SID] Fimbristyl
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REFERENCES: Coville 1890; Bush 1905
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RHYNCHOSPORA/CYPERACEAE 621 Lipocar
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RHYNCHOSPORA/CYPERACEAE 623 10. Per
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RHYNCHOSPORA/CYPERACEAE 625 34. Cul
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Lipocarpha aristulata [GWO] Lipocar
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SCHOENOPLECTUS/CYPERACEAE 641 exten
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Schoenoplectus pungens [MAS] Schoen
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SCLERIA/CYPERACEAE 649 5. Lowest la
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SCLERIA/CYPERACEAE 651 with a pair
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ERIOCAULON/ERIOCAULACEAE 657 1. Pet
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ERIOCAULON/ERIOCAULACEAE 659 bracts
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� Dioscorea bulbifera [HEB] ILLUS
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HYACINTHACEAE 663 orange zone towar
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Scleria pauciflora ILLUSTRATED FLOR
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� � ILLUSTRATED FLORA OF EAST T
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SCHOENOLIRION/HYACINTHACEAE 669 som
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HYDROCHARITACEAE 671 AA small (116
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HYDRILLIA/HYDROCHARITACEAE 673 marg
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OTTELIA Pers. DUCK-LETTUCE OTTELIA/
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� Vallisneria americana [LAM] Ott
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IRIDACEAE 683 inflorescences with 1
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BELAMCANDA Adans. LEOPARD-LILY, BLA
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IRIS/IRIDACEAE 691 6. Cauline leaf
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Iris brevicaulis [YAT] ILLUSTRATED
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SISYRINCHIUM/IRIDACEAE 697 3. Inflo
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Sisyrinchium ensigerum [CUR] Sisyri
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JUNCUS/JUNCACEAE 703 mainly in wet
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JUNCUS/JUNCACEAE 705 yellowish, lea
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Juncus acuminatus [MAS] Juncus anth
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Juncus dichotomus [GWO] Juncus effu
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Juncus polycephalus [GWO] Juncus re
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var. validus-● var. facinatus-�
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LEMNA/LEMNACEAE 721 Pineywoods and
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SPIRODELA/LEMNACEAE 723 Lemna turio
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NOTE 3 VEINS � TOP VIEW Lemna min
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LILIACEAE 729 The lily family (even
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ERYTHRONIUM/LILIACEAE 731 17. Tepal
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Wolffia columbiana [GWO, MIB] Wolff
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LIMNOCHARITACEAE 735 1. Perianth wh
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THALIA/MARANTACEAE 737 pollen prese
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MELANTHIACEAE 739 AA small family (
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Lilium philippinense [GCH] SEED FOL
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VERATRUM/MELANTHIACEAE 745 circumsc
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Zigadenus glaberrimus [GLE] Schoeno
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ALETRIS/NARTHECIACEAE 749 Tofieldia
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DASYLIRION Zucc. SOTOL, SHAGGY-LILY
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FLOWER Aletris aurea [CO1] Aletris
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ORCHIDACEAE 755 45 occur in East TX
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CALOPOGON/ORCHIDACEAE 757 5-25 flow
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CLEISTES/ORCHIDACEAE 759 long; lip
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CYPRIPEDIUM/ORCHIDACEAE 761 while t
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Stenanthium gramineum Toxicoscordio
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Calopogon tuberosus [FNA] Corallorh
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HEXALECTRIS/ORCHIDACEAE 767 rain. A
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Habenaria quinqueseta [LUN] Habenar
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PLATANTHERA/ORCHIDACEAE 771 PLATANT
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PLATANTHERA/ORCHIDACEAE 773 Platant
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SPIRANTHES/ORCHIDACEAE 779 mm wide,
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SPIRANTHES/ORCHIDACEAE 781 12. Lip
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Platanthera lacera [YAT] Pogonia op
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SPIRANTHES/ORCHIDACEAE 789 narrowes
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TRIPHORA/ORCHIDACEAE 791 an example
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LEAF collar rhizome blade ligule sh
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POACEAE 797 Dissection is usually n
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POACEAE 799 rower and often differe
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POACEAE 801 6. Lemmas awned, the tw
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POACEAE 803 30. Lemma veins none to
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POACEAE 805 13. Inflorescences narr
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POACEAE 807 12. Lemma awnless; infl
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POACEAE 809 inflorescence solitary,
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POACEAE 811 29. Spikelets loosely a
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POACEAE 813 20. Basal leaves usuall
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AEGILOPS/POACEAE 815 23. Spikelets
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AGROSTIS/POACEAE 817 Agrostis hyema
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ANDROPOGON/POACEAE 821 colonizing e
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ANTHENANTIA/POACEAE 825 to be the e
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ARISTIDA/POACEAE 829 9. Lemma thick
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ARISTIDA/POACEAE 831 BRIT), Newton
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Anthoxanthum aristatum [BB2] Anthox
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AXONOPUS/POACEAE 839 duced to the U
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Avena fatua [HI1] Arundo donax [LYN
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BOTHRIOCHLOA/POACEAE 843 var. barbi
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Aristida purpurea (all vars.) ILLUS
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Axonopus compressus [HI1] Axonopus
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BOUTELOUA/POACEAE 849 state grass b
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Bothriochloa ischaemum var. songari
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Bouteloua pectinata [BOT, HEA] Bout
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BROMUS/POACEAE 857 8. Lemmas and gl
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BUCHLOE/POACEAE 861 ticularly in th
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CHASMANTHIUM/POACEAE 865 [C. caroli
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CHLORIS/POACEAE 871 East TX; mainly
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Chasmanthium sessiliflorum [USB] Ch
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CORTADERIA/POACEAE 875 of the inflo
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DANTHONIA/POACEAE 881 Dactylocteniu
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Cynodon dactylon [REE] Cynosurus ec
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DICHANTHELIUM/POACEAE 885 AA genus
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DICHANTHELIUM/POACEAE 887 20. Inter
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DICHANTHELIUM/POACEAE 889 26. Leaf
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Danthonia spicata [USB] Desmazeria
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DICHANTHELIUM/POACEAE 897 and Harri
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Dichanthelium acuminatum (all vars.
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DICHANTHELIUM/POACEAE 905 that this
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DIGITARIA/POACEAE 909 2. Main axis
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Dichanthelium sphaerocarpon [HI] IL
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DIGITARIA/POACEAE 913 with papilla-
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Digitaria bicornis [HEB] ILLUSTRATE
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Digitaria insularis [HI1] Digitaria
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EHRHARTA/POACEAE 921 spikelets (2.5
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ELIONURUS/POACEAE 923 Since this co
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Echinochloa crus-gali [REE] Echinoc
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ENTEROPOGON Nees UMBRELLA GRASS ENT
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ERAGROSTIS/POACEAE 931 8. Infloresc
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ERAGROSTIS/POACEAE 933 12. Pedicels
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Elymus repens [HI1] Elymus smithii
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Eragrostis japonica [USB] ILLUSTRAT
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ERIONEURON/POACEAE 945 Shaw and Sme
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� � � ILLUSTRATED FLORA OF EA
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FESTUCA/POACEAE 951 rachilla glabro
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GYMNOPOGON/POACEAE 955 Glyceria sep
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HILARIA/POACEAE 957 REFERENCES: Eme
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ILLUSTRATED FLORA OF EAST TEXAS Fes
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KOELERIA/POACEAE 963 Andropogoneae,
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Heteropogon melanocarpus [HI1] Hord
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LEPTOCHLOA/POACEAE 967 Leersia oryz
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ILLUSTRATED FLORA OF EAST TEXAS Koe
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ILLUSTRATED FLORA OF EAST TEXAS Hor
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LOLIUM/POACEAE 973 Charmet et al. 1
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Leptochloa nealleyi [USB] Lolium ri
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MICROSTEGIUM/POACEAE 977 AA variabl
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ILLUSTRATED FLORA OF EAST TEXAS Lep
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Lolium temulentum [USB] Melica muti
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MUHLENBERGIA/POACEAE 983 10. Lemma
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MUHLENBERGIA/POACEAE 985 and Prairi
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ILLUSTRATED FLORA OF EAST TEXAS Mon
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ILLUSTRATED FLORA OF EAST TEXAS Mic
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 993 fact that gener
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PANICUM/POACEAE 995 17. Spikelets 2
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Oryza sativa [HI1] Panicum anceps [
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 1001 Panicum hallii
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Panicum trichoides [HI2] ILLUSTRATE
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PASPALUM/POACEAE 1009 5. Spikelets
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PASPALUM/POACEAE 1011 27. Axils of
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Panicum virgatum [RCA] Paspalum acu
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PASPALUM/POACEAE 1019 loam soils; w
Page 713 and 714: ILLUSTRATED FLORA OF EAST TEXAS 102
Page 717 and 718: PENNISETUM/POACEAE 1025 unit as in
Page 719 and 720: Paspalum repens [USB] ILLUSTRATED F
Page 721 and 722: PHALARIS/POACEAE 1029 landscapes (B
Page 723 and 724: Paspalum pubiflorum (both vars.) IL
Page 725 and 726: Pennisetum purpureum [HUT] Penniset
Page 727 and 728: POA/POACEAE 1035 around awn base; l
Page 729 and 730: POA/POACEAE 1037 long. Forests, flo
Page 737 and 738: SACCHARUM/POACEAE 1045 1. Awn of le
Page 739 and 740: SCHEDONNARDUS/POACEAE 1047 lower gl
Page 745 and 746: SETARIA/POACEAE 1053 Paspalidium (W
Page 747 and 748: SETARIA/POACEAE 1055 14. Leaf blade
Page 749 and 750: Schizachyrium tenerum [HI1, USB] Sc
Page 755 and 756: SORGHUM/POACEAE 1063 2000) indicate
Page 757 and 758: SPARTINA/POACEAE 1065 Americas it [
Page 759 and 760: Sorghastrum elliottii [HI1] Sorghas
Page 763: SPOROBOLUS/POACEAE 1071 7. Lemma ap
Page 767 and 768: STEINCHISMA/POACEAE 1075 Sporobolus
Page 769 and 770: Sphenopholis obtusata (both vars.)
Page 771 and 772: Sporobolus neglectus [HI1] Sporobol
Page 773 and 774: TRICHONEURA/POACEAE 1081 guish it f
Page 775 and 776: Stenotaphrum secundatum [USB] Trich
Page 781 and 782: TRISETUM/POACEAE 1089 low areas; ne
Page 783 and 784: UROCHLOA/POACEAE 1091 AA C4 genus o
Page 787 and 788: Tridens eragrostoides Tridens flavu
Page 789 and 790: VULPIA/POACEAE 1097 2. Lemmas (exce
Page 795 and 796: ZIZANIOPSIS/POACEAE 1103 (1999) (po
Page 799 and 800: EICHHORNIA/PONTEDERIACEAE 1107 REFE
Page 801 and 802: Zoysia japonica [HOL] Zoysia pacifi
Page 803 and 804: POTAMOGETON/POTAMOGETONACEAE 1111 1
Page 805 and 806: POTAMOGETON/POTAMOGETONACEAE 1113 P
Page 811 and 812: SMILAX/SMILACACEAE 1119 times woody
Page 813 and 814: SMILAX/SMILACACEAE 1121 30(-60) mm
Page 815 and 816:
Stuckenia pectinata [MAS] ILLUSTRAT
Page 817 and 818:
Page 819 and 820:
THEMIDACEAE 1127 cover for various
Page 821 and 822:
Smilax rotundifolia [JEM] ILLUSTRAT
Page 823 and 824:
TRILLIUM/TRILLIACEAE 1131 Schilling
Page 825 and 826:
TRILLIUM/TRILLIACEAE 1133 taxonomic
Page 827 and 828:
Triantha racemosa [GLE] Trillium lu
Page 829 and 830:
TYPHA/TYPHACEAE 1137 REFERENCES: Wi
Page 831 and 832:
XYRIS/XYRIDACEAE 1139 cone-like spi
Page 833 and 834:
XYRIS/XYRIDACEAE 1141 11. Plant bas
Page 835 and 836:
Sparganium americanum Typha latifol
Page 837 and 838:
Typha domingensis[ BT3] Typha latif
Page 839 and 840:
Xyris drummondii [SID] Xyris fimbri
Page 841 and 842:
Xyris smalliana [SID] ILLUSTRATED F
Page 843:
Xyris caroliniana Xyris difformis (
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keys to the vascular plants of east texas - Botanical Research ...

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