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DICHANTHELIUM/POACEAE 905<br />

that this species appears related <strong>to</strong> D. oligosan<strong>the</strong>s but differs in having wider leaf blades, larger<br />

spikelets, and bearded culm nodes.<br />

Dichan<strong>the</strong>lium scabriusculum (Elliott) Gould & Clark, (somewhat rough), TALL SWAMP PANIC<br />

GRASS, WOOLLY ROSETTE GRASS. Culms usually 90–150 cm tall, <strong>the</strong> nodes nei<strong>the</strong>r swollen nor<br />

bearded, glabrous or only slightly hairy, <strong>of</strong>ten with a glandular band below <strong>the</strong> node; leaf<br />

sheaths glabrous or with some pubescence, but not velvety pubescent, narrowed at <strong>to</strong>p, mottled<br />

with pale spots; leaf blades 7–15 mm wide, glabrous or with some pubescence on lower surface,<br />

but not velvety pubescent; spikelets 2.2–2.9 mm long. Low, wet, <strong>of</strong>ten shaded sites, sandy soils;<br />

scattered in East TX in Jasper (Ajilvsgi 5921, BRIT), Hardin, Leon, Tyler (TAES—annotated by S.<br />

Hatch), Anderson (SBSC), Austin, Bowie, and Robertson (Turner et al. 2003) cos.; e U.S. from NY<br />

s <strong>to</strong> FL w <strong>to</strong> TX. May(–Jul), also fall. [Panicum scabriusculum Elliott] This species is somewhat<br />

similar <strong>to</strong> D. scoparium but has leaf sheaths glabrous or at l<strong>east</strong> without velvety pubescence<br />

and nodes not bearded. According <strong>to</strong> Freckmann and Lelong (2003a), D. scabriusculum possibly<br />

hybridizes with D. clandestinum, D. dicho<strong>to</strong>mum, and D. aciculare.<br />

Dichan<strong>the</strong>lium scoparium (Lam.) Gould, (broom-like), VELVET ROSETTE GRASS, VELVETY PANIC<br />

GRASS. Culms coarse, 80–155 cm tall, <strong>of</strong>ten conspicuously s<strong>to</strong>ut (usually 2–5 mm thick), <strong>the</strong><br />

nodes <strong>of</strong>ten swollen, usually conspicuously bearded with a glabrous, glandular band below <strong>the</strong><br />

beard; lower culm internodes and lower leaf sheaths and blades usually with velvety pubescence;<br />

leaf sheaths narrowed at <strong>to</strong>p; leaf blades (7–)9–25(–30) mm wide; spikelets 2.2–2.8 mm<br />

long, <strong>of</strong>ten purplish at base. Sandy woods and low areas, <strong>of</strong>ten in disturbed sites; widespread in<br />

East TX, mainly in Pineywoods and Post Oak Savannah; also n Gulf Prairies and Marshes; e U.S.<br />

from NY s <strong>to</strong> FL w <strong>to</strong> KS and TX. May–Jun, also fall. [Panicum scoparium Lam.] If not for <strong>the</strong><br />

velvety pubescence, this species would resemble D. scabriusculum (Gould 1975b).<br />

Dichan<strong>the</strong>lium scoparium possibly hybridizes with D. acuminatum and D. dicho<strong>to</strong>mum<br />

(Freckmann & Lelong 2003a).<br />

Dichan<strong>the</strong>lium sphaerocarpon (Elliott) Gould, (spherical-fruited), ROUND-SEED ROSETTE GRASS,<br />

ROUND-SEED PANIC, ROUND-SEED DICHANTHELIUM, ROUND-FRUIT PANIC GRASS. Culms 20–50 cm<br />

tall; ligule minute or apparently absent; blades <strong>of</strong> larger culm leaves 6–13(–14) mm wide, usually<br />

cordate at base, slightly auricled-clasping, ciliate only near <strong>the</strong> base, <strong>the</strong> margins white and<br />

cartilaginous; inflorescence axis glabrous; spikelets 1.4–1.8(–2) mm long. Sandy soils, shaded or<br />

open areas; widespread in e 1/2 <strong>of</strong> TX, scattered fur<strong>the</strong>r w; se Canada (Ont.) and widespread in<br />

e U.S. w <strong>to</strong> KS and TX. Late Mar–Jun, again late summer–fall. [Panicum sphaerocarpon Elliott]<br />

The <strong>plants</strong> are similar <strong>to</strong> glabrous forms <strong>of</strong> D. acuminatum var. lindheimeri but differ in having<br />

a minute or absent ligule. This species also sometimes superficially resembles D. strigosum. It<br />

can, however, be distinguished by <strong>the</strong> glabrous inflorescence axis and leaf blades ciliate only<br />

near <strong>the</strong> base (versus inflorescence axis usually with spreading hairs and leaf blades ciliate <strong>to</strong><br />

beyond <strong>the</strong> middle in D. strigosum). Dichan<strong>the</strong>lium sphaerocarpon hybridizes with several o<strong>the</strong>r<br />

species, including D. acuminatum, D. laxiflorum, and D. polyan<strong>the</strong>s (Gould & Clark 1978;<br />

Freckmann & Lelong 2003a). This species has sometimes been treated as including D. polyan<strong>the</strong>s<br />

as a variety (var. isophyllum or var. polyan<strong>the</strong>s); Gould (1975b) indicated that this and D.<br />

polyan<strong>the</strong>s, which he treated as D. sphaerocarpon var. polyan<strong>the</strong>s, seem <strong>to</strong> completely intergrade.<br />

Dichan<strong>the</strong>lium strigosum (Muhl.) Freckmann, (strigose, with stiff bristles), ROUGH-HAIR RO-<br />

SETTE GRASS, CUSHION-TUFTED PANIC GRASS. Culms ca. 45 cm or less tall, not branched above<br />

base; leaves mostly basal (only 2–4 leaves on culm); leaf sheaths glabrous or ascending pilose;<br />

leaf blades 3–8 mm wide, ciliate <strong>to</strong> beyond <strong>the</strong> middle; inflorescence axis with spreading hairs;<br />

spikelets 1.1–2.1 mm long, <strong>of</strong>ten purplish or brownish purple; glumes ca. 1/3–1/2 as long as<br />

spikelet. Moist pinelands, bogs, o<strong>the</strong>r low areas, sandy soils. The infraspecific taxa <strong>of</strong> this species<br />

have been variously treated as varieties, subspecies, or species (e.g., Turner et al. 2003). Until<br />

a thorough revision <strong>of</strong> <strong>the</strong> group is carried out, we are recognizing <strong>the</strong> infraspecific taxa at <strong>the</strong>

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