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keys to the vascular plants of east texas - Botanical Research ...

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742 MELANTHIACEAE/STENANTHIUM<br />

South TX Plains; in <strong>the</strong> U.S. this species is known only from TX. Sep–Oct (rarely in spring). This<br />

species has in TX long gone under <strong>the</strong> name S. drummondii A. Gray (e.g., Correll & Johns<strong>to</strong>n<br />

1970), which is now considered an illegitimate name (TROPICOS 2003). Frame (2002) treated<br />

this species as S. ghiesbreghtii Greenm. (also known from Mexico), but based on molecular<br />

work in preparation (W. Zomlefer, pers. comm.), <strong>the</strong> U.S. populations likely warrant recognition<br />

as a separate species. According <strong>to</strong> Frame (2002), in <strong>the</strong> U.S. this species “grows at or near sea<br />

level and almost exclusively on <strong>the</strong> Reyosa gravel formation <strong>of</strong> <strong>the</strong> Texas Gulf prairie. … In <strong>the</strong><br />

late 1930s and early 1940s, it was used in a series <strong>of</strong> transplantation experiments conducted by<br />

<strong>the</strong> Texas Agricultural Experiment Stations. This has led <strong>to</strong> some confusion with regard <strong>to</strong> its<br />

natural distribution.” Until <strong>the</strong> nomenclature <strong>of</strong> this species is clarified, we are following <strong>the</strong> recent<br />

Flora <strong>of</strong> North America treatment (Frame 2002) in using <strong>the</strong> name S. ghiesbreghtii for <strong>the</strong><br />

GREEN-LILY.<br />

Schoenocaulon texanum Scheele, (<strong>of</strong> Texas), TEXAS SABADILLA, TEXAS FEATHER-SHANK. Plant <strong>to</strong> 55<br />

cm tall; leaves <strong>to</strong> 60 cm long, mostly 4(–6) mm or less wide; tepals 2.3–4 mm long, greenish<br />

white; capsules 10–15(–19) mm long; 2n = 16 (Frame 2002). Limes<strong>to</strong>ne soils, rocky grasslands<br />

and openings in juniper-oak woodlands; Bell, Hays, Travis (BRIT), Comal (ASTC), Bexar (Turner<br />

et al. 2003), and Williamson (Balcones Canyonlands Nat. Wildlife Refuge, C. Sex<strong>to</strong>n, pers.<br />

comm.) cos.; w margin <strong>of</strong> East TX s <strong>to</strong> South TX Plains and w through Edwards Plateau <strong>to</strong><br />

Trans-Pecos; NM and TX. Mar–Jul (–rarely later after unseasonable rainfall). According <strong>to</strong><br />

Frame (2002), “The morphology <strong>of</strong> Schoenocaulon texanum is peculiarly dependent upon annual<br />

rainfall. Plants <strong>of</strong> small stature but average bulb size can be readily distinguished as <strong>the</strong><br />

products <strong>of</strong> one or several seasons <strong>of</strong> lower than normal rainfall. The hallmarks <strong>of</strong> such <strong>plants</strong><br />

are congested racemes, few flowers, <strong>the</strong>se <strong>of</strong>ten appearing wi<strong>the</strong>red and brown, and ovaries that<br />

do not mature or have perhaps only one small locule ripening <strong>to</strong> produce a few seeds.” m/297<br />

STENANTHIUM (A. Gray) Kunth FEATHER-BELLS, FEATHER-FLEECE<br />

Perennial, subscapose, glabrous herbs from a slender bulb; leaves crowded near base; inflorescence<br />

a raceme or panicle; flowers perfect and/or staminate; tepals 6, all � alike, greenish white<br />

<strong>to</strong> creamy white, cream, or yellowish, separate <strong>to</strong> base, not clawed, glandless or with an obscure<br />

gland at base <strong>of</strong> each; stamens 6; ovary usually superior or partly inferior; fruit a capsule.<br />

AA genus <strong>of</strong> 2 or 3 species <strong>of</strong> <strong>the</strong> e U.S. (Zomlefer & Judd 2002; Zomlefer & Smith 2002). Recent<br />

chromosomal, molecular, and morphological studies (Zomlefer et al. 2001; Zomlefer &<br />

Judd 2002; Zomlefer & Smith 2002) have necessitated significant changes in generic circumscription<br />

<strong>of</strong> Stenanthium. Several species previously placed in Stenanthium, as well as several<br />

formerly in Zigadenus, have now been shown <strong>to</strong> belong in <strong>the</strong> genus Anticlea (Zomlefer et al.<br />

2001; Zomlefer & Judd 2002; Zomlefer & Smith 2002). Stenanthium gramineum (<strong>the</strong> type species<br />

<strong>of</strong> <strong>the</strong> genus) has been shown <strong>to</strong> be most closely related <strong>to</strong> taxa (S. densum, S. leimanthoides<br />

if recognized) previously treated in Zigadenus. This is a clear example <strong>of</strong> new research requiring<br />

scientific names <strong>to</strong> be changed (despite some nomenclatural confusion). See fur<strong>the</strong>r discussion<br />

and references under Zigadenus.<br />

All species <strong>of</strong> Stenanthium have a chromosome number <strong>of</strong> 2n = 20 (Zomlefer & Smith 2002).<br />

(Greek: stenos, narrow, and anthos, flower, in reference <strong>to</strong> <strong>the</strong> narrow inflorescences or tepals)<br />

REFERENCES: Fernald 1946b; Zomlefer et al. 2001; Utech 2002b; Zomlefer & Judd 2002;<br />

Zomlefer & Smith 2002.<br />

1. Tepals 3–5 mm long, each with an obscure gland near <strong>the</strong> base; inflorescence a raceme or panicle<br />

5 cm or less wide; pedicels 10–20 mm long; capsules 10–20 mm long ________________________ S. densum<br />

1. Tepals (4–)5–10 mm long, without glands; inflorescence a panicle <strong>of</strong>ten > 5 cm wide; pedicels<br />

4–6 mm long; capsules 6–10 mm long _____________________________________________ S. gramineum

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