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ALETRIS/NARTHECIACEAE 749<br />

T<strong>of</strong>ieldiaceae, which are in a clade including Alismataceae and Araceae, near <strong>the</strong> base <strong>of</strong> <strong>the</strong><br />

monocots (Chase et al. 2000; Fuse & Tamura 2000). For a detailed discussion <strong>of</strong> <strong>the</strong> groups formerly<br />

treated as Liliaceae in <strong>the</strong> broad sense, see <strong>the</strong> family synopsis <strong>of</strong> <strong>the</strong> Liliaceae (here<br />

treated in a restricted sense) on page 726. Family name from Nar<strong>the</strong>cium, BOG ASPHODEL, a genus<br />

<strong>of</strong> 4–8 species <strong>of</strong> temperate North America, e Asia, and w Europe. (Name an anagram <strong>of</strong><br />

An<strong>the</strong>ricum L.—Zomlefer 1997c, a genus previously treated in a broadly defined Liliaceae but<br />

now in <strong>the</strong> An<strong>the</strong>ricaceae—Mabberley 1997) (subclass Liliidae—Cronquist; order Dioscoreales—<br />

APG II)<br />

FAMILY RECOGNITION IN THE FIELD: scapose perennials with small white <strong>to</strong> yellow flowers in a<br />

spike-like raceme, <strong>the</strong> tepals free only at <strong>the</strong>ir tips and wrinkled and roughened externally and<br />

appearing mealy.<br />

REFERENCES: Dahlgren et al. 1985; Goldblatt 1995; Zomlefer 1997a, 1997c, 1999; Reveal &<br />

Zomlefer 1998; Tamura 1998c; Caddick et al. 2000, 2002a, 2002b; Fuse & Tamura 2000; Davis et<br />

al. 2004.<br />

ALETRIS L. STAR-GRASS, COLIC-ROOT<br />

Perennial, glabrous herbs from a short thick rhizome; leaves mostly in a basal rosette, sessile,<br />

linear-lanceolate <strong>to</strong> elliptic, flat; flowers in a spike-like raceme terminating a nearly naked<br />

scape (a few small remote bracts can be present); flowers perfect, each subtended by 2 subulate,<br />

unequal bracts; perianth tubular <strong>to</strong> campanulate, wrinkled and roughened externally (appearing<br />

mealy or farinose), <strong>the</strong> 6 segments (= tepals) free only at <strong>the</strong> apex (lobes); stamens 6, attached<br />

<strong>to</strong> perianth just below perianth lobes; style 1, with 3 stigmas; ovary superior <strong>to</strong> partly<br />

inferior; fruit a beaked capsule.<br />

AA genus <strong>of</strong> ca. 15–25 species <strong>of</strong> North America, <strong>the</strong> West Indies (Bahamas), and e Asia<br />

(Zomlefer 1997c; Sullivan 2002), including several cultivated as ornamentals. Some authors<br />

(e.g., Wen 1999) consider this genus an example <strong>of</strong> an e Asia-e North America disjunction; this<br />

distribution pattern is discussed under <strong>the</strong> genus Brachyelytrum (Poaceae). The rhizomes <strong>of</strong><br />

some species (particularly A. farinosa) are rich in steroidal sapogenins and have been used medicinally,<br />

giving rise <strong>to</strong> such common names as COLIC-ROOT, AGUE-ROOT, BREAST-ROOT, BACK-<br />

ACHE-ROOT, and RHEUMATISM-ROOT (Cheatham et al. 1995; Zomlefer 1997c). (Greek: aletris, a female<br />

slave who ground corn, referring <strong>to</strong> <strong>the</strong> apparent mealiness <strong>of</strong> <strong>the</strong> perianth)<br />

REFERENCES: Browne 1956, 1961; Sullivan 2002.<br />

1. Perianth yellow <strong>to</strong> orange-yellow, ca. 6–7 mm long, <strong>the</strong> lobes ovate <strong>to</strong> triangular ________________ A. aurea<br />

1. Perianth white <strong>to</strong> creamy-white, usually (7–)8–10 mm long, <strong>the</strong> lobes lanceolate <strong>to</strong> lanceoblong<br />

_________________________________________________________________________ A. farinosa<br />

Aletris aurea Walter, (golden), YELLOW STAR-GRASS, YELLOW COLIC-ROOT, GOLDEN COLIC-ROOT.<br />

Leaves flat, lanceolate, 2–8(–12) cm long, 10–20 mm wide, bright yellow green; scape 20–80 cm<br />

or more tall; pedicels 3 mm or less long; perianth yellow <strong>to</strong> orange-yellow (can fade upon drying),<br />

roughened on <strong>the</strong> outside; beak <strong>of</strong> fruit gradually narrowed. Savannahs, boggy areas, pine<br />

barrens; Pineywoods and Post Oak Savannah; also n Gulf Prairies and Marshes; se U.S. from MD<br />

s <strong>to</strong> FL w <strong>to</strong> AR, OK, and TX. May–Jul. m/273<br />

Aletris farinosa L., (mealy, powdery), UNICORN-ROOT, WHITE-TUBE STAR-GRASS, WHITE STAR-<br />

GRASS, SOUTHERN COLIC-ROOT, WHITE COLIC-ROOT. Leaves 4–15(–20) cm long, (5–)10–20(–26)<br />

mm wide, dark green; scape 40–100 cm tall; perianth white <strong>to</strong> creamy white, roughened on <strong>the</strong><br />

outside; beak <strong>of</strong> fruit abruptly narrowed. Sandy soils, gravels, and peats; tentatively included<br />

based on citations for East TX by Correll and Johns<strong>to</strong>n (1970) and for <strong>the</strong> ne corner <strong>of</strong> East TX<br />

(without county specified) by Sullivan (2002); no county distribution map is provided; se<br />

Canada and widespread in <strong>the</strong> e U.S. w <strong>to</strong> IL, AR, and TX. Late spring–mid-summer. m/273

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