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keys to the vascular plants of east texas - Botanical Research ...

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ASPARAGACEAE<br />

475<br />

spreading from cultivation in Austin (Travis Co.) and has escaped in Bandera Co. (Edwards Plateau);<br />

in <strong>the</strong> U.S. known only from TX, but widespread from Mexico <strong>to</strong> Central America.<br />

Spring–Fall. [S. exul (O.F. Cook) L.H. Bailey, S. texana (O.F. Cook) Becc.] While rare in TX, S.<br />

mexicana is “one <strong>of</strong> <strong>the</strong> most common palms <strong>of</strong> lowland tropical Mexico” (Zona 1990). The species<br />

is <strong>of</strong>ten cultivated in <strong>the</strong> s part <strong>of</strong> TX (but can survive winters much fur<strong>the</strong>r north—e.g., Austin<br />

and even Dallas—Lockett 2003). Because <strong>the</strong> wood was useful in wharf building (resistant <strong>to</strong><br />

<strong>the</strong> shipworm, which destroys wood in warm salt water) and <strong>the</strong> <strong>plants</strong> prized as ornamentals<br />

(Lockett & Read 1990; Lockett 2003), it is likely that most native populations have been harvested,<br />

and that this species was much more common in presettlement times. (TOES 1993: IV) �<br />

Sabal minor (Jacq.) Pers., (smaller), DWARF PALMETTO, BUSH PALMETTO, DWARF PALM, BLUE PALM,<br />

BLUE STEM, SWAMP PALM, BLUE PALMETTO, SWAMP PALMETTO, LATANIA, LATANIER. Plant <strong>to</strong> ca. 2 m tall<br />

(rarely taller—<strong>to</strong> nearly 5 m), usually acaulescent with a pithy crown or rarely with a short<br />

trunk (10–20 cm in diam.); leaf blades � suborbicular in outline, <strong>to</strong> ca. 1.6 m wide. Usually in<br />

stream bot<strong>to</strong>ms, floodplains, riverbanks, and swamps (but curiously also known from dry,<br />

open, upland habitats in parts <strong>of</strong> <strong>the</strong> Edwards Plateau—e.g., Lockett 1991); Pineywoods and Gulf<br />

Prairies and Marshes w along larger rivers <strong>to</strong> Dallas, Kaufman (BRIT), and McLennan (BAYLU)<br />

cos. in <strong>the</strong> Blackland Prairie and Gonzales (Turner et al. 2003; Palmet<strong>to</strong> State Park), Van Zandt,<br />

and Wood (BRIT) cos. in <strong>the</strong> Post Oak Savannah, also Hays and Travis cos. on w margin <strong>of</strong> East<br />

TX w <strong>to</strong> Gillespie Co. (BRIT) on <strong>the</strong> Edwards Plateau and naturalized in Tarrant Co. (R.<br />

O’Kennon, pers. obs.); se U.S. from NC s <strong>to</strong> FL w <strong>to</strong> OK and TX; recently a population was discovered<br />

in n Mexico, ca. 225 miles sw <strong>of</strong> <strong>the</strong> nearest known population (Goldman 1999); this<br />

area <strong>of</strong> Mexico is known <strong>to</strong> share a number <strong>of</strong> floristic components with <strong>the</strong> e U.S. (Dressler<br />

1954; Martin & Harrell 1957). Mainly spring–summer, but nearly year round in lower Rio<br />

Grande Valley (Davis in Lockett & Read 1990). [S. louisiana (Darby) Bomhard] While S. minor is<br />

typically acaulescent, populations <strong>of</strong> trunked individuals are known, especially in <strong>the</strong> w portion<br />

<strong>of</strong> <strong>the</strong> species range (particularly in LA and TX); stems <strong>of</strong> 1–1.5 m in height are not uncommon<br />

(Ramp & Thien 1995). Such populations have long caused taxonomic confusion (e.g., <strong>the</strong><br />

name S. louisiana). Bailey (1961) indicated that “There appear <strong>to</strong> be no botanical characters<br />

separating <strong>the</strong> two extremes except such dimensions <strong>of</strong> parts as are associated with stature.”<br />

Recent electrophoretic studies support <strong>the</strong> inclusion <strong>of</strong> <strong>the</strong> trunked populations in S. minor<br />

(Ramp & Thien 1995). A Brazoria Co. population (<strong>of</strong> larger trunked individuals up <strong>to</strong> 27 feet<br />

tall) may be hybids between S. minor and S. mexicana (Lockett 1991, 2003). According <strong>to</strong> Zona<br />

(2000), “Fur<strong>the</strong>r research is needed <strong>to</strong> test this hypo<strong>the</strong>sis.” Part <strong>of</strong> <strong>the</strong> hybrid population is now<br />

protected in a satellite unit <strong>of</strong> <strong>the</strong> San Bernard National Wildlife Refuge (Lockett 2003). m/296<br />

ASPARAGACEAE Juss.<br />

ASPARAGUS FAMILY<br />

AA small family <strong>of</strong> 2 genera and ca. 305 species (Judd 2001). In addition <strong>to</strong> Asparagus, <strong>the</strong> family<br />

includes only Hemiphylacus, a small Mexican group <strong>of</strong> 5 species (Hernández 1995). While previously<br />

<strong>of</strong>ten placed in <strong>the</strong> Asphodelaceae or Hyacinthaceae, recent evidence (primarily molecular<br />

data) suggests that Hemiphylacus be included in <strong>the</strong> Asparagaceae (e.g., Rudall et al. 1997, 1998b;<br />

Judd 2001). Members <strong>of</strong> <strong>the</strong> Asparagaceae are typically adapted <strong>to</strong> dry conditions. According <strong>to</strong><br />

Kubitzki and Rudall (1998), “Most species are found in regions with semiarid <strong>to</strong> arid and Mediterranean-type<br />

climate, and extreme xeromorphic adaptations are common.” Many authorities have<br />

put <strong>the</strong> taxa here treated as Asparagaceae in a broadly defined and clearly polyphyletic (but<br />

practical) Liliaceae (e.g., Correll & Johns<strong>to</strong>n 1970; Cronquist 1988; Diggs et al. 1999), based on superficial<br />

similarities in flower structure <strong>to</strong> <strong>the</strong> genus Lilium. O<strong>the</strong>rs (e.g., Mabberley 1997) have<br />

recognized <strong>the</strong> Asparagaceae but have treated it in a broad sense (6 genera). However, based on<br />

phylogenetic analyses, we are following Judd (2001) in recognizing <strong>the</strong> Asparagaceae as a mono-

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