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726 LILIACEAE fronds originate); roots absent; flowers 1(–2) per frond, in cavity beside mid-line of upper surface of frond, but plants seldom flowering; in quiet waters. AA genus of 10 species (Landolt 2000), mostly of tropical and warm areas of the Americas and Africa. The following key is based on non-flowering material because “Wolffiella fronds change shape drastically when they are fertile …” (Godfrey & Wooten 1979). (Name a diminutive of Wolffia) REFERENCES: Landolt 1984; Crawford et al. 1997. 1. Fronds (4–)6–15(–20) times longer than wide, pointed at apex (= tip, the end opposite the end where the daughter fronds originate), narrowly sabre-shaped, long-tapered from a slightly broader base to the pointed apex; angle of vegetative reproductive pouch 25°–50° __________________ W. gladiata 1. Fronds 1.5-8 times longer than wide, rounded or pointed at apex, tongue-shaped to ribbon-like or ovate, the sides � parallel for much of their length or short-tapered; angle of vegetative reproductive pouch 45°–120°. 2. Area of air spaces in tissue near base of frond usually no longer than wide, occupying basal 1/3 to 1/2 of frond; fronds 1.5–4 times longer than wide, usually curved downward at apex; angle of vegetative reproductive pouch 70°–120° ___________________________________ W. lingulata 2. Area of air spaces in tissue near base of frond usually longer than wide, occupying basal 2/3 or more of frond; fronds 3–8 times longer than wide, usually flat; angle of vegetative reproductive pouch 45°–90° __________________________________________________________ W. oblonga Wolffiella gladiata (Hegelm.) Hegelm., (sword-like), SWORD BOG-MAT. Fronds narrowly sabreshaped, 4–9 mm long, ca. 0.5–1.4 mm wide at base, distinctly long-tapered from a slightly broader base to a pointed apex, often asymmetric near the apex with a curvature or an indentation on one side, rarely solitary to usually 2–many (to 20 or more) attached, forming star-like groups; area of air spaces within frond much longer than wide; seldom flowering. Scattered in East TX; also Gulf Prairies and Marshes.; throughout most of the e U.S. w to IL and TX, also WA. [Wolffia gladiata Hegelm., Wolffia gladiata var. floridana Donn. Sm., Wolffiella floridana (Donn. Sm.) C.H. Thomps.] Wolffiella lingulata (Hegelm.) Hegelm., (strap-shaped), TONGUE-SHAPED BOG-MAT. Fronds tongue-shaped to widely so or ovate, 3–10 mm long, rounded at apex, � symmetric to strongly curved, solitary to 2(–4) attached; seldom flowering. Harrison Co. (Evans & Brown 2002 based on Nixon 16,133, ASTC); also Brazoria Co. (Turner et al. 2003); also Landolt (2000) mapped it for the Gulf Prairies and Marshes just s of the Big Thicket; CA, FL, LA, MS, and TX. [Wolffia lingulata Hegelm.] Wolffiella oblonga (Phil.) Hegelm., (elliptical with blunt ends), SABER BOG-MAT. Fronds narrowly tongue-shaped to ribbon-like, 1.2–7.5 mm long, rounded or pointed at apex, � symmetric near the apex, solitary or 2–8 attached, forming star-like groups; seldom flowering. Jasper Co. (BRIT, Diggs & George, s.n.—B.A. Steinhagen Lake, mixed collection with W. gladiata), also range map in Landolt (2000) showing occurrence in s part of the Pineywoods (without specific county); also Gulf Prairies and Marshes (Landolt 2000); CA, FL, LA, MS, and TX. [Lemna oblonga Phil.] While not officially designated as such (e.g., TOES 1993; Carr 2002d; Poole et al. 2002), given its limited distribution in the state, we consider this species to be of conservation concern in TX. � LILIACEAE Juss. LILY FAMILY Perennial herbs from a bulb; leaves alternate or whorled along the stem or basal; flowers 1–few in a terminal umbel-like cluster or solitary at end of a naked scape; flowers conspicuous; tepals 6, distinct, all petaloid; stamens 6; ovary superior, 3-carpellate; style 1; fruit a capsule.
ILLUSTRATED FLORA OF EAST TEXAS 727 Landoltia punctata Lemna aequinoctialis Lemna minuta Lemna obscura Lemma perpusilla Lemna turionifera Lemna valdiviana Spirodela polyrhiza Wolffia brasiliensis Wolffia columbiana Wolffiella gladiata Wolffiella lingulata
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KEYS TO THE VASCULAR PLANTS 309 KEY
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KEY TO FERNS AND SIMILAR PLANTS 311
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Key to Families of Gymnosperms KEY
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20. Flowers in a single, small (4-4
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38. Stamens 6 per flower. 39. Peria
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57. Leaves reduced to scales only;
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FERNS AND SIMILAR PLANTS (PTERIDOPH
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ISOETES/ISOETACEAE 323 DIVISION LYC
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Psilotum nudum [KNO] Isoetes butler
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LYCOPODIELLA/LYCOPODIACEAE 327 Beit
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SELAGINELLA/SELAGINELLACEAE 329 Pse
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ILLUSTRATED FLORA OF EAST TEXAS 331
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Palhinhaea cernua [CLU] Selaginella
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leaf blade petiole rhizome root sca
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ASPLENIUM L. SPLEENWORT ASPLENIUM/A
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WOODWARDIA/BLECHNACEAE 339 1. Plant
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MEGASPORE Woodwardia areolata [LUN]
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DRYOPTERIDACEAE 343 AThe family as
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Anemia mexicana Azolla caroliniana
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DRYOPTERIS/DRYOPTERIDACEAE 347 Carr
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WOODSIA/DRYOPTERIDACEAE 351 Tectari
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MARSILEA/MARSILEACEAE 353 etation (
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BOTRYCHIUM/OPHIOGLOSSACEAE 357 REFE
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Nephrolepis exaltata Onoclea sensib
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OPHIOGLOSSUM/OPHIOGLOSSACEAE 361 Op
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PTERIDACEAE 365 and pous or podium,
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Botrychium virginianum Ophioglossum
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CERATOPTERIS/PTERIDACEAE 371 Astrol
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Argyrochosma dealbata [LUN] Astrole
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Cheilanthes feei [LUN] Cheilanthes
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Astrolepis integerrima Astrolepis s
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SALVINIACEAE 379 fern, derived from
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SALVINIA/SALVINIACEAE 381 NM, and N
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Pellaea ovata Pellaea wrightiana IL
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Thelypteris dentata [SM3] STERILE F
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GYMNOSPERMS EPHEDRACEAE 389 AThe te
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DIVISION PINOPHYTA CONIFERS CUPRESS
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JUNIPERUS/CUPRESSACEAE 393 allergen
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Ephedra antisyphilitica [LUN] Junip
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PINUS/PINACEAE 397 clusters at the
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� � � � � � Pinus echin
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Ephedra antisyphilitica Juniperus a
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MAGNOLIOPHYTA 403 consumed in large
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CLASS MONOCOTYLEDONAE MONOCOTYLEDON
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AGAVACEAE 407 produced in North Ame
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HESPERALOE/AGAVACEAE 409 for their
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Acorus calamus [GLE] Hesperaloe eng
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YUCCA/AGAVACEAE 413 snakebite (Verh
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7. Leaves straight or nearly so, la
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Manfreda virginica subsp. lata [HEA
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Manfreda maculosa Manfreda sileri I
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ALISMA/ALISMATACEAE 423 1. Pistils
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Yucca rupicola [LYN] Yucca treculea
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SAGITTARIA/ALISMATACEAE 427 long te
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Echinodorus tenellus [FLN, RHO] Sag
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Yucca treculeana ILLUSTRATED FLORA
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ALLIUM/ALLIACEAE 433 pounds are ant
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ALLIUM/ALLIACEAE 435 Allium ampelop
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STAMENS ILLUSTRATED FLORA OF EAST T
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CAPSULE Allium canadense var. mobil
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var. fraseri-▲ var. hyacinthoides
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AMARYLLIDACEAE 443 and edible (Baye
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COOPERIA/AMARYLLIDACEAE 445 REFEREN
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BULBILS CAPSULE Allium sativum [ANO
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HABRANTHUS/AMARYLLIDACEAE 449 obser
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CAPSULES SEED Habranthus tubispathu
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LYCORIS/AMARYLLIDACEAE 455 usually
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Hymenocallis liriosme [HEA] Leucoju
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Narcissus jonquilla [NIC] Narcissus
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ANTHERICACEAE 461 2000), Anderson,
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ARACEAE 463 hemiepiphytes, and aqua
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ARISAEMA/ARACEAE 465 has been disag
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ORONTIUM/ARACEAE 469 down the insid
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Echeandia flavescens [MCV] Colocasi
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ARECACEAE 473 oped rhizomes present
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ASPARAGACEAE 475 spreading from cul
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BROMELIACEAE 477 jointed pedicels;
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Xanthosoma sagittifolium [BA1] Saba
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APTERIA/BURMANNIACEAE 481 FAMILY RE
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S. minor � S. mexicana-� ILLUST
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UVULARIA/COLCHICACEAE 485 blurred,
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ANTHER CAPSULE PISTIL Apteria aphyl
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CALLISIA/COMMELINACEAE 489 the Haem
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COMMELINA/COMMELINACEAE 491 4. Spat
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SINGLE FLOWER INFLORESCENCE Callisi
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TRADESCANTIA/COMMELINACEAE 495 & Tv
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FRUITING INFLORESCENCE Murdannia nu
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Tradescantia paludosa [AND] Tradesc
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CYPERACEAE 507 become split by grow
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CYPERACEAE 509 10. Scales of spikel
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BULBOSTYLIS/CYPERACEAE 511 1. Spike
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CAREX/CYPERACEAE 517 The following
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CAREX/CYPERACEAE 519 of leaf sheath
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CAREX/CYPERACEAE 521 37. Perigynia
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CAREX/CYPERACEAE 523 58. Achene sho
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CAREX/CYPERACEAE 525 85. Larger per
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Carex abscondita [MAC] Carex albica
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CAREX/CYPERACEAE 531 persists in li
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Carex decomposita [MAC] Carex digit
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Carex triangularis [MAC] Carex tetr
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CYPERUS/CYPERACEAE 559 1974; Baijan
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CYPERUS/CYPERACEAE 561 21. Leaf bla
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CYPERUS/CYPERACEAE 563 fresh; achen
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CYPERUS/CYPERACEAE 565 tively more
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CYPERUS/CYPERACEAE 573 Cyperus invo
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Cyperus esculentus [REE] Cyperus fi
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CYPERUS/CYPERACEAE 577 around the s
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Cyperus iria [GWO] Cyperus lanceola
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Cyperus surinamensis [BT2] Cyperus
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ELEOCHARIS/CYPERACEAE 587 5. Culms
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ELEOCHARIS/CYPERACEAE 589 27. Spike
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ELEOCHARIS/CYPERACEAE 591 43. Scale
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ELEOCHARIS/CYPERACEAE 593 compressa
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ELEOCHARIS/CYPERACEAE 601 ing mats;
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FIMBRISTYLIS Vahl FIMBRY FIMBRISTYL
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Fimbristylis annua [SID] Fimbristyl
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REFERENCES: Coville 1890; Bush 1905
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RHYNCHOSPORA/CYPERACEAE 621 Lipocar
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RHYNCHOSPORA/CYPERACEAE 623 10. Per
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RHYNCHOSPORA/CYPERACEAE 625 34. Cul
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Lipocarpha aristulata [GWO] Lipocar
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SCHOENOPLECTUS/CYPERACEAE 641 exten
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Schoenoplectus pungens [MAS] Schoen
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SCLERIA/CYPERACEAE 649 5. Lowest la
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SCLERIA/CYPERACEAE 651 with a pair
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ERIOCAULON/ERIOCAULACEAE 657 1. Pet
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ERIOCAULON/ERIOCAULACEAE 659 bracts
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� Dioscorea bulbifera [HEB] ILLUS
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HYACINTHACEAE 663 orange zone towar
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Scleria pauciflora ILLUSTRATED FLOR
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� � ILLUSTRATED FLORA OF EAST T
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SCHOENOLIRION/HYACINTHACEAE 669 som
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HYDROCHARITACEAE 671 AA small (116
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HYDRILLIA/HYDROCHARITACEAE 673 marg
Page 367 and 368: ILLUSTRATED FLORA OF EAST TEXAS 675
Page 369 and 370: OTTELIA Pers. DUCK-LETTUCE OTTELIA/
Page 373 and 374: � Vallisneria americana [LAM] Ott
Page 375 and 376: IRIDACEAE 683 inflorescences with 1
Page 377 and 378: BELAMCANDA Adans. LEOPARD-LILY, BLA
Page 383 and 384: IRIS/IRIDACEAE 691 6. Cauline leaf
Page 385 and 386: Iris brevicaulis [YAT] ILLUSTRATED
Page 389 and 390: SISYRINCHIUM/IRIDACEAE 697 3. Inflo
Page 393 and 394: Sisyrinchium ensigerum [CUR] Sisyri
Page 395 and 396: JUNCUS/JUNCACEAE 703 mainly in wet
Page 397 and 398: JUNCUS/JUNCACEAE 705 yellowish, lea
Page 401 and 402: Juncus acuminatus [MAS] Juncus anth
Page 403 and 404: Juncus dichotomus [GWO] Juncus effu
Page 407 and 408: Juncus polycephalus [GWO] Juncus re
Page 409 and 410: var. validus-● var. facinatus-�
Page 413 and 414: LEMNA/LEMNACEAE 721 Pineywoods and
Page 415 and 416: SPIRODELA/LEMNACEAE 723 Lemna turio
Page 417: NOTE 3 VEINS � TOP VIEW Lemna min
Page 421 and 422: LILIACEAE 729 The lily family (even
Page 423 and 424: ERYTHRONIUM/LILIACEAE 731 17. Tepal
Page 425 and 426: Wolffia columbiana [GWO, MIB] Wolff
Page 427 and 428: LIMNOCHARITACEAE 735 1. Perianth wh
Page 429 and 430: THALIA/MARANTACEAE 737 pollen prese
Page 431 and 432: MELANTHIACEAE 739 AA small family (
Page 433 and 434: Lilium philippinense [GCH] SEED FOL
Page 437 and 438: VERATRUM/MELANTHIACEAE 745 circumsc
Page 439 and 440: Zigadenus glaberrimus [GLE] Schoeno
Page 441 and 442: ALETRIS/NARTHECIACEAE 749 Tofieldia
Page 443 and 444: DASYLIRION Zucc. SOTOL, SHAGGY-LILY
Page 445 and 446: FLOWER Aletris aurea [CO1] Aletris
Page 447 and 448: ORCHIDACEAE 755 45 occur in East TX
Page 449 and 450: CALOPOGON/ORCHIDACEAE 757 5-25 flow
Page 451 and 452: CLEISTES/ORCHIDACEAE 759 long; lip
Page 453 and 454: CYPRIPEDIUM/ORCHIDACEAE 761 while t
Page 455 and 456: Stenanthium gramineum Toxicoscordio
Page 457 and 458: Calopogon tuberosus [FNA] Corallorh
Page 459 and 460: HEXALECTRIS/ORCHIDACEAE 767 rain. A
Page 461 and 462: Habenaria quinqueseta [LUN] Habenar
Page 463 and 464: PLATANTHERA/ORCHIDACEAE 771 PLATANT
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SPIRANTHES/ORCHIDACEAE 779 mm wide,
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SPIRANTHES/ORCHIDACEAE 781 12. Lip
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Platanthera lacera [YAT] Pogonia op
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SPIRANTHES/ORCHIDACEAE 789 narrowes
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TRIPHORA/ORCHIDACEAE 791 an example
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LEAF collar rhizome blade ligule sh
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POACEAE 797 Dissection is usually n
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POACEAE 799 rower and often differe
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POACEAE 801 6. Lemmas awned, the tw
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POACEAE 803 30. Lemma veins none to
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POACEAE 805 13. Inflorescences narr
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POACEAE 807 12. Lemma awnless; infl
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POACEAE 809 inflorescence solitary,
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POACEAE 811 29. Spikelets loosely a
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POACEAE 813 20. Basal leaves usuall
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AEGILOPS/POACEAE 815 23. Spikelets
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AGROSTIS/POACEAE 817 Agrostis hyema
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ANDROPOGON/POACEAE 821 colonizing e
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ANTHENANTIA/POACEAE 825 to be the e
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ARISTIDA/POACEAE 829 9. Lemma thick
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ARISTIDA/POACEAE 831 BRIT), Newton
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Anthoxanthum aristatum [BB2] Anthox
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AXONOPUS/POACEAE 839 duced to the U
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Avena fatua [HI1] Arundo donax [LYN
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BOTHRIOCHLOA/POACEAE 843 var. barbi
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Aristida purpurea (all vars.) ILLUS
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Axonopus compressus [HI1] Axonopus
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BOUTELOUA/POACEAE 849 state grass b
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Bothriochloa ischaemum var. songari
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Bouteloua pectinata [BOT, HEA] Bout
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BROMUS/POACEAE 857 8. Lemmas and gl
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BUCHLOE/POACEAE 861 ticularly in th
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CHASMANTHIUM/POACEAE 865 [C. caroli
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CHLORIS/POACEAE 871 East TX; mainly
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Chasmanthium sessiliflorum [USB] Ch
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CORTADERIA/POACEAE 875 of the inflo
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DANTHONIA/POACEAE 881 Dactylocteniu
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Cynodon dactylon [REE] Cynosurus ec
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DICHANTHELIUM/POACEAE 885 AA genus
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DICHANTHELIUM/POACEAE 887 20. Inter
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DICHANTHELIUM/POACEAE 889 26. Leaf
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Danthonia spicata [USB] Desmazeria
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DICHANTHELIUM/POACEAE 897 and Harri
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Dichanthelium acuminatum (all vars.
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DICHANTHELIUM/POACEAE 905 that this
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DIGITARIA/POACEAE 909 2. Main axis
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Dichanthelium sphaerocarpon [HI] IL
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DIGITARIA/POACEAE 913 with papilla-
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Digitaria bicornis [HEB] ILLUSTRATE
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Digitaria insularis [HI1] Digitaria
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EHRHARTA/POACEAE 921 spikelets (2.5
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ELIONURUS/POACEAE 923 Since this co
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Echinochloa crus-gali [REE] Echinoc
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ENTEROPOGON Nees UMBRELLA GRASS ENT
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ERAGROSTIS/POACEAE 931 8. Infloresc
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ERAGROSTIS/POACEAE 933 12. Pedicels
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Elymus repens [HI1] Elymus smithii
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Eragrostis japonica [USB] ILLUSTRAT
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ERIONEURON/POACEAE 945 Shaw and Sme
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� � � ILLUSTRATED FLORA OF EA
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FESTUCA/POACEAE 951 rachilla glabro
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GYMNOPOGON/POACEAE 955 Glyceria sep
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HILARIA/POACEAE 957 REFERENCES: Eme
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ILLUSTRATED FLORA OF EAST TEXAS Fes
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KOELERIA/POACEAE 963 Andropogoneae,
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Heteropogon melanocarpus [HI1] Hord
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LEPTOCHLOA/POACEAE 967 Leersia oryz
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ILLUSTRATED FLORA OF EAST TEXAS Koe
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ILLUSTRATED FLORA OF EAST TEXAS Hor
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LOLIUM/POACEAE 973 Charmet et al. 1
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Leptochloa nealleyi [USB] Lolium ri
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MICROSTEGIUM/POACEAE 977 AA variabl
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ILLUSTRATED FLORA OF EAST TEXAS Lep
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Lolium temulentum [USB] Melica muti
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MUHLENBERGIA/POACEAE 983 10. Lemma
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MUHLENBERGIA/POACEAE 985 and Prairi
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ILLUSTRATED FLORA OF EAST TEXAS Mon
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ILLUSTRATED FLORA OF EAST TEXAS Mic
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 993 fact that gener
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PANICUM/POACEAE 995 17. Spikelets 2
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Oryza sativa [HI1] Panicum anceps [
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 1001 Panicum hallii
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Panicum trichoides [HI2] ILLUSTRATE
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PASPALUM/POACEAE 1009 5. Spikelets
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PASPALUM/POACEAE 1011 27. Axils of
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Panicum virgatum [RCA] Paspalum acu
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PASPALUM/POACEAE 1019 loam soils; w
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PENNISETUM/POACEAE 1025 unit as in
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Paspalum repens [USB] ILLUSTRATED F
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PHALARIS/POACEAE 1029 landscapes (B
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Paspalum pubiflorum (both vars.) IL
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Pennisetum purpureum [HUT] Penniset
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POA/POACEAE 1035 around awn base; l
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POA/POACEAE 1037 long. Forests, flo
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SACCHARUM/POACEAE 1045 1. Awn of le
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SCHEDONNARDUS/POACEAE 1047 lower gl
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SETARIA/POACEAE 1053 Paspalidium (W
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SETARIA/POACEAE 1055 14. Leaf blade
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Schizachyrium tenerum [HI1, USB] Sc
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SORGHUM/POACEAE 1063 2000) indicate
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SPARTINA/POACEAE 1065 Americas it [
Page 759 and 760:
Sorghastrum elliottii [HI1] Sorghas
Page 761 and 762:
Page 763 and 764:
SPOROBOLUS/POACEAE 1071 7. Lemma ap
Page 765 and 766:
Page 767 and 768:
STEINCHISMA/POACEAE 1075 Sporobolus
Page 769 and 770:
Sphenopholis obtusata (both vars.)
Page 771 and 772:
Sporobolus neglectus [HI1] Sporobol
Page 773 and 774:
TRICHONEURA/POACEAE 1081 guish it f
Page 775 and 776:
Stenotaphrum secundatum [USB] Trich
Page 777 and 778:
Page 779 and 780:
Page 781 and 782:
TRISETUM/POACEAE 1089 low areas; ne
Page 783 and 784:
UROCHLOA/POACEAE 1091 AA C4 genus o
Page 785 and 786:
Page 787 and 788:
Tridens eragrostoides Tridens flavu
Page 789 and 790:
VULPIA/POACEAE 1097 2. Lemmas (exce
Page 791 and 792:
Page 793 and 794:
Page 795 and 796:
ZIZANIOPSIS/POACEAE 1103 (1999) (po
Page 797 and 798:
Page 799 and 800:
EICHHORNIA/PONTEDERIACEAE 1107 REFE
Page 801 and 802:
Zoysia japonica [HOL] Zoysia pacifi
Page 803 and 804:
POTAMOGETON/POTAMOGETONACEAE 1111 1
Page 805 and 806:
POTAMOGETON/POTAMOGETONACEAE 1113 P
Page 807 and 808:
Page 809 and 810:
Page 811 and 812:
SMILAX/SMILACACEAE 1119 times woody
Page 813 and 814:
SMILAX/SMILACACEAE 1121 30(-60) mm
Page 815 and 816:
Stuckenia pectinata [MAS] ILLUSTRAT
Page 817 and 818:
Page 819 and 820:
THEMIDACEAE 1127 cover for various
Page 821 and 822:
Smilax rotundifolia [JEM] ILLUSTRAT
Page 823 and 824:
TRILLIUM/TRILLIACEAE 1131 Schilling
Page 825 and 826:
TRILLIUM/TRILLIACEAE 1133 taxonomic
Page 827 and 828:
Triantha racemosa [GLE] Trillium lu
Page 829 and 830:
TYPHA/TYPHACEAE 1137 REFERENCES: Wi
Page 831 and 832:
XYRIS/XYRIDACEAE 1139 cone-like spi
Page 833 and 834:
XYRIS/XYRIDACEAE 1141 11. Plant bas
Page 835 and 836:
Sparganium americanum Typha latifol
Page 837 and 838:
Typha domingensis[ BT3] Typha latif
Page 839 and 840:
Xyris drummondii [SID] Xyris fimbri
Page 841 and 842:
Xyris smalliana [SID] ILLUSTRATED F
Page 843:
Xyris caroliniana Xyris difformis (
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keys to the vascular plants of east texas - Botanical Research ...

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