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1016 POACEAE/PASPALUM<br />

Guadalupe Co. (Turner et al. 2003) near sw margin <strong>of</strong> East TX; mainly Gulf Prairies and<br />

Marshes and South TX Plains; in <strong>the</strong> U.S. known only from TX; also Latin America. Jun–Nov.<br />

Paspalum laeve Michx., (smooth), FIELD PASPALUM, FIELD CROWN GRASS. Tufted perennial without<br />

rhizomes or only shortly rhizoma<strong>to</strong>us; culms <strong>to</strong> ca. 100(–120) cm tall; leaf sheaths glabrous <strong>to</strong><br />

pilose; leaf blades sometimes marginally crisped; inflorescence branches (2–)3–6(–8), (2–)4–<br />

10(–12) cm long, <strong>the</strong> branch axes � unwinged; spikelets 2.4–3.4 mm long, solitary, in 2 rows;<br />

lower glume absent; upper glume and lemma <strong>of</strong> sterile floret glabrous. Prairies, open woods,<br />

disturbed or moist areas, <strong>of</strong>ten on sand; Pineywoods and Post Oak Savannah; also n Gulf Prairies<br />

and Marshes; e U.S. from MA s <strong>to</strong> FL w <strong>to</strong> KS and TX. Jul–Oct. While some authorities (e.g.,<br />

Kartesz 1999; Turner et al. 2003; Allen & Hall 2003) do not recognize varieties in this species,<br />

until a detailed study <strong>of</strong> <strong>the</strong> group is carried out, we are following Gould (1975b), Jones et al.<br />

(1997), Yatskievych (1999), and Hatch (2002) in recognizing <strong>the</strong> 3 somewhat overlapping varieties—not<br />

all individuals can be assigned with certainty. The maps in Turner et al. (2003) and <strong>the</strong><br />

one presented here do not distinguish varieties.<br />

1. Spikelets 2.7–3.2 mm wide, ca. as wide as long and circular in outline or nearly so _____________ var. circulare<br />

1. Spikelets 2–2.5 mm wide, conspicuously longer than wide, broadly ovate <strong>to</strong> broadly obovate in<br />

outline.<br />

2. Lower leaf sheaths and lower leaf blades glabrous or with sparse hairs ______________________ var. laeve<br />

2. Lower leaf sheaths and <strong>of</strong>ten lower leaf blades moderately <strong>to</strong> densely hairy _______________ var. pilosum<br />

var. circulare (Nash) Fernald, (circular), ROUND-SEED PASPALUM. Lamar, Liberty, Marion, San<br />

Augustine, and Upshur (BRIT) cos. [P. circulare Nash]<br />

var. laeve, FIELD PASPALUM, SMOOTH PASPALUM. Anderson, Frees<strong>to</strong>ne, Henderson, Hous<strong>to</strong>n, Jefferson,<br />

Morris, Panola, Rusk, Shelby, Titus, Upshur (BRIT), and Lamar (Carr 1994) cos. This is <strong>the</strong><br />

most common variety in East TX.<br />

var. pilosum Scribn., (with long s<strong>of</strong>t hairs), HAIR-LEAF PASPALUM. Panola and Titus (BRIT) cos. [P.<br />

longipilum Nash]<br />

Paspalum langei (E. Fourn.) Nash, (for Johann Martin Christian Lange, 1818–1898, Danish botanist),<br />

RUSTY-SEED PASPALUM, RUSTY-SEED CROWN GRASS, LANGE’S PASPALUM. Cespi<strong>to</strong>se perennial<br />

without rhizomes; culms 30–125 cm tall; ligule ca. 0.6–2 mm long; leaf sheaths and blades glabrous<br />

or with minute pubescence; leaf blades (4–)7–18 mm broad, <strong>of</strong>ten conspicuously crisped<br />

marginally; inflorescence branches (1–)2–5, usually 4–10(–13) cm long, <strong>the</strong> branch axes �<br />

unwinged; spikelets usually 2.2–2.7 mm long, usually paired, appearing in 2 or 4 rows; lower<br />

glume present on some or all spikelets, reduced; upper glume and lemma <strong>of</strong> sterile floret pubescent,<br />

usually with brownish glandular blotches. Shaded ditches, stream bot<strong>to</strong>ms, moist<br />

woods; s part <strong>of</strong> East TX; also Gulf Prairies and Marshes, South TX Plains, e Edwards Plateau,<br />

and disjunct <strong>to</strong> Tarrant Co. (BRIT) in Cross Timbers and Prairies; FL, LA, OK, and TX. Apr–Nov.<br />

[Dimorphostachys langei E. Fourn.]<br />

Paspalum lividum Trin., (lead-colored, bluish gray), LONG-TOM, PULL-AND-BE-DAMNED. Tufted perennial;<br />

culms decumbent <strong>to</strong> s<strong>to</strong>loniferous basally, <strong>to</strong> 70 cm tall but <strong>to</strong> over 1.5 m long; ligule (1–<br />

)2.2–4.7 mm long; inflorescence branches usually 3–7(–11), (1–)1.5–4(–5) cm long, <strong>the</strong> branch<br />

axes winged; spikelets 2.2–2.6 mm long, typically paired, usually in 4 rows; lower glume absent;<br />

upper glume and lemma <strong>of</strong> sterile floret glabrous. Ditches, wet places, muddy coastal areas,<br />

fresh or brackish water; Bexar, Colorado, Harris, Jasper, Jefferson, Orange, Waller, and Wilson<br />

(Turner et al. 2003) cos. on s margin <strong>of</strong> East TX; mainly Gulf Prairies and Marshes and South<br />

TX Plains, also Burnet and Brown (Turner et al. 2003) cos. near w margin <strong>of</strong> Cross Timbers and<br />

Prairies; FL, LA, and TX. May–Nov. This species is important for cattle grazing along <strong>the</strong> Gulf<br />

Coast (Hatch et al. 1999).

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