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CYPERACEAE<br />

509<br />

10. Scales <strong>of</strong> spikelets with short bristle-tip or bristle-tip absent; perianth <strong>of</strong> 1 thin<br />

perianth scale, or <strong>of</strong> bristles, or perianth absent.<br />

11. Spikelets (actually small spikelet-like spikes or heads <strong>of</strong> spirally arranged<br />

single-flowered spikelets) 1–5(–8) mm long, sessile; each scale enclosing<br />

a single, thin, inconspicuous perianth-like bracteole; <strong>plants</strong> very small, <strong>to</strong><br />

only 15(–20) cm tall; scales with 2 or 3 prominent veins (use strong hand<br />

lens) _________________________________________________________ Lipocarpha<br />

11. Spikelets 2.5–20 mm long, sessile or on distinct pedicels; each scale enclosing<br />

a perianth <strong>of</strong> bristles or perianth absent; <strong>plants</strong> small <strong>to</strong> very large;<br />

scales with only 1 prominent vein (<strong>the</strong> midrib).<br />

12. Spikelets with a single fertile (= achene-producing) floret subtended<br />

by ca. 2–5 empty scales (<strong>the</strong>se lack achenes but can have an aborted<br />

pistil or stamens); leaf margins scaberulous (= only slightly roughened,<br />

almost smooth <strong>to</strong> <strong>the</strong> <strong>to</strong>uch) <strong>to</strong> dangerously spinulose-serrulate (=<br />

saw-<strong>to</strong>o<strong>the</strong>d); inflorescences with numerous spikelets, <strong>the</strong>se usually<br />

in groups <strong>of</strong> 2–10 at <strong>the</strong> ends <strong>of</strong> short branches; perianth absent;<br />

species rare in East TX __________________________________________ Cladium<br />

12. Spikelets not as above, with 1–many fertile florets, each spikelet subtended<br />

by at most 1 empty scale; leaf margins various; inflorescences<br />

not as above; perianth absent or <strong>of</strong> bristles; including species common<br />

and widespread in East TX.<br />

13. Inflorescences usually ei<strong>the</strong>r 1-sided or widely spreading or drooping;<br />

perianth <strong>of</strong> bristles (except in Isolepis and Schoenoplectus<br />

saximontanus); <strong>plants</strong> small <strong>to</strong> very large (<strong>to</strong> 5 m tall) (segregates<br />

<strong>of</strong> Scirpus in <strong>the</strong> broad sense).<br />

14. Inflorescences with 1(–2) spreading <strong>to</strong> erect modified leaf (involucral<br />

bract) appearing like a continuation <strong>of</strong> <strong>the</strong> culm, <strong>the</strong><br />

inflorescences thus appearing lateral; culms with 1–3 leaves<br />

near base OR without blade-bearing leaves.<br />

15. Plants rhizoma<strong>to</strong>us perennials 30–300(–500) cm tall; culms<br />

2–20 mm thick near base; achenes smooth; perianth<br />

bristles present; inflorescences with 1–150 or more<br />

spikelets ____________________________________ Schoenoplectus<br />

15. Plants tufted annuals or perennials 2–65 cm tall; culms<br />

0.2–1.5 mm thick near base; achenes minutely papillose<br />

(sometimes obscurely so) or with prominent, transverse,<br />

wavy ridges; perianth bristles absent.<br />

16. Achenes minutely papillose (this can sometimes be<br />

obscured by a whitish wax-like layer); spikelets 2–10<br />

mm long; inflorescences with 1–3 spikelets; <strong>plants</strong> 2–<br />

25(–40) cm tall ____________________________________ Isolepis<br />

16. Achenes with prominent, transverse, wavy ridges;<br />

spikelets 5–20 mm long; inflorescences with1–10(–20)<br />

spikelets; <strong>plants</strong> 9–65 cm tall _________________ Schoenoplectus<br />

14. Inflorescences with 2 or more well-developed leaf-like involucral<br />

bracts, <strong>the</strong> inflorescences appearing terminal; culms with<br />

well-developed leaves.<br />

17. Spikelets small, 3–10 mm long, 2–4 mm wide, very numerous;<br />

scales <strong>of</strong> spikelets glabrous, not apically notched, awnless<br />

or nearly so; achenes < 1.5 mm long; culms mostly<br />

obscurely triangular ___________________________________ Scirpus

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