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ERIOCAULON/ERIOCAULACEAE<br />

659<br />

bracts; outer involucral bracts narrowly ovate <strong>to</strong> lanceolate, 2–4 mm long, acuminate, strawcolored,<br />

with clavate white hairs apically; surface <strong>of</strong> receptacle villous; receptacular bracts narrowly<br />

acute <strong>to</strong> acuminate, sometimes visibly exserted; sepals 2–3 mm long, yellowish white,<br />

with clavate white hairs on keel and apex. Meadows, swamps, pond margins, bogs; widespread<br />

in appropriate habitats in <strong>the</strong> Pineywoods and Post Oak Savannah; also n Gulf Prairies and<br />

Marshes; e U.S. from NY s <strong>to</strong> FL w <strong>to</strong> OK and TX. May–Sep. This is <strong>the</strong> most robust <strong>of</strong> <strong>the</strong> East<br />

TX pipeworts. m/286<br />

Eriocaulon koernickianum Van Heurck & Müll. Arg., (for Friedrich August Körnicke, 1828–<br />

1908, German botanist), SMALL-HEAD PIPEWORT, GULF PIPEWORT, DWARF PIPEWORT. Plant annual<br />

or weakly perennial, diminutive, <strong>the</strong> leaves 1–5 cm long, <strong>the</strong> mature scapes 5–8 cm long; sheath<br />

<strong>of</strong> scapes about <strong>the</strong> length <strong>of</strong> <strong>the</strong> leaves; mature heads subglobose or short-oblong, 3–4 mm<br />

broad, dark gray or gray-green except for <strong>the</strong> pale “rims” <strong>of</strong> white ciliate perianth parts and<br />

bracts and pale outer bracts; outer involucral bracts broadly oblong <strong>to</strong> suborbicular, reflexed at<br />

maturity, 1–1.25 mm long, rounded, straw-colored, translucent, not hidden by perianth parts<br />

and receptacular bracts; surface <strong>of</strong> receptacle glabrous; receptacular bracts acute <strong>to</strong> obtuse; sepals<br />

ca. 1 mm long, gray-translucent, glabrous or with scattered hairs on <strong>the</strong> back apically. Upland<br />

seeps, bogs, permanently wet seepage slopes or marsh/bog edges that are not overshadowed<br />

by taller vegetation, <strong>of</strong>ten on sphagnum mats, acidic sands <strong>to</strong> sandy silts, <strong>of</strong>ten associated<br />

with sands<strong>to</strong>ne or o<strong>the</strong>r rocky outcrops; Anderson, Brazos, Frees<strong>to</strong>ne (BRIT), Polk, Tyler (Kral<br />

1966a), Henderson, Limes<strong>to</strong>ne (MacRoberts & MacRoberts 1999), and Leon (Carr 2001) cos. in<br />

<strong>the</strong> s portions <strong>of</strong> <strong>the</strong> Post Oak Savannah and Pineywoods (MacRoberts & MacRoberts (1999)<br />

could not confirm <strong>the</strong> Polk and Tyler county records); also Gillespie Co. (BRIT) at Enchanted<br />

Rock State Natural Area in <strong>the</strong> e Edwards Plateau; AR, GA, OK, and TX. Spring–early fall. This<br />

rare species was cited by Kral (2000b) as being <strong>of</strong> conservation concern, has declined in <strong>the</strong> recent<br />

past, and is a Federal Candidate Species (Watson et al. 1994). Its conservation biology was<br />

discussed by Watson et al. (1994)—a number <strong>of</strong> fac<strong>to</strong>rs appear <strong>to</strong> be contributing <strong>to</strong> its rareness,<br />

including no vegetative reproduction, low seed set, genetic homogeneity, poor competitive ability,<br />

annual or weak perennial life his<strong>to</strong>ry, and a restricted habitat. (TOES 1993: V; RARE 2002a:<br />

G2S1SOC) �<br />

Eriocaulon texense Körn., (<strong>of</strong> Texas), TEXAS PIPEWORT, PIPEWORT. In habit resembling a diminutive<br />

E. compressum (Kral 2000b); leaves 1–5 cm long; mature scapes 5–30 cm long, mature head<br />

usually hemispherical, 5–10 mm broad, gray except for <strong>the</strong> white hairs <strong>of</strong> <strong>the</strong> receptacular<br />

bracts and perianth parts and <strong>the</strong> straw-colored outer involucral bracts, s<strong>of</strong>t; outer involucral<br />

bracts suborbicular <strong>to</strong> broadly ovate, ca. 1.5 mm long, rounded <strong>to</strong> acute, at maturity usually<br />

hidden by <strong>the</strong> florets (but evident earlier); surface <strong>of</strong> receptacle with numerous hairs;<br />

receptacular bracts acute; sepals ca. 1.5 mm long, dark gray apically, with clavate white hairs on<br />

keel and sometimes on margins apically. Wet acidic areas, sphagnous bogs (<strong>of</strong>ten with Sarracenia—PITCHERPLANT);<br />

widespread in s part <strong>of</strong> Pineywoods and Post Oak Savannah w <strong>to</strong> Henderson,<br />

Milam (TAES), Austin, and Frees<strong>to</strong>ne (Turner et al. 2003) cos.; se U.S. from NC s <strong>to</strong> FL w <strong>to</strong><br />

TX. Moldenke (1961) indicated Cory recorded <strong>the</strong> species from <strong>the</strong> Blackland Prairie. Apr–Jun.<br />

Eriocaulon aquaticum (Hill) Druce, (aquatic), SEVEN-ANGLE PIPEWORT, WHITE-BUTTONS, DUCK-<br />

GRASS, [E. septangulare With.], was reported for TX in vegetational areas 1 and 4 by Hatch et al.<br />

(1990). However, Kral (1979b; 2000b) indicated it occurs in Canada and <strong>the</strong> n U.S. s only <strong>to</strong> Alabama;<br />

R. Kral (pers. comm.) confirmed that this species does not occur in TX. According <strong>to</strong> Kral,<br />

“Early reports <strong>of</strong> E. aquaticum (E. septangulare) were based on lack <strong>of</strong> understanding <strong>of</strong> <strong>the</strong><br />

similar-looking E. texense.” Eriocaulon aquaticum can be distinguished by <strong>the</strong> heads which are<br />

small (4–5 mm wide) and � dark in appearance due <strong>to</strong> usually gray <strong>to</strong> almost black<br />

receptacular bracts and perianth parts (only tips <strong>of</strong> perianth parts and receptacular bracts<br />

have whitish hairs); also <strong>the</strong> receptacles are glabrous.

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