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1092 POACEAE/UROCHLOA 8. Upper glume without distinct cross veins; upper glume and lemma of lower floret usually puberulent; lemma of lower floret with 5 veins ___________________________ U. ramosa Urochloa ciliatissima (Buckley) R.D. Webster, (very fringed with hairs), FRINGED SIGNAL GRASS, FRINGED LIVER-SEED GRASS, SANDHILL GRASS. Perennial 10–50 cm tall; culms erect or usually decumbent below; inflorescence with few, short, spreading or ascending branches and relatively few spikelets; spikelets (3–)3.5–4.5 mm long; lower glume glabrous, ca. 3/4 as long as spikelet; upper glume densely long-hairy; lemma of sterile floret glabrous except for pilose margins. Rocky or sandy open sites; widespread in TX but more commonly in the s and w parts of the state; AR, NM, OK, and TX. May–Sep. [Brachiaria ciliatissima (Buckley) Chase] Urochloa fusca (Sw.) B.F. Hansen & Wunderlin, (brown, dusky), HURRAH GRASS, BROWNTOP, BROWN-TOP SIGNAL GRASS, FIELD GRASS, BROWN-TOP LIVER-SEED GRASS. Annual with spreading to erect, branching culms 10–85 cm long; inflorescence with primary branches appressed or erect-spreading, unbranched or sometimes rebranched, the branches and pedicels often with some long whitish hairs; spikelets 2.4–3.5 mm long; lower glume 1/4–1/3 as long as spikelet; glumes and lemma of lower floret glabrous, often brownish. Dried-up pond or stream margins, roadsides, disturbed sites; throughout much of TX but less common in the Pineywoods and Panhandle; s U.S. from FL w to AZ. May–Oct. [Brachiaria fasciculata (Sw.) Parodi, Brachiaria fasciculata var. reticulata (Torr.) Vickery, Panicum fasciculatum Sw. var. reticulatum (Torr.) Beal, Panicum reticulatum Torr., U. fasciculata var. reticulata (Torr.) R.D. Webster—invalid] The complicated nomenclatural history of this species was discussed by Wunderlin and Hansen (2001). Urochloa maxima (Jacq.) R.D. Webster, (largest), GUINEA GRASS, GUINEA LIVER-SEED GRASS. Rhizomatous perennial; culms (0.6–)1–2.5 m tall; inflorescence an open panicle 20–50(–65) cm long, the branches at lowest node in whorls, the primary branches with secondary and tertiary branches (the plant thus superficially very much resembling Panicum); spikelets 2.7–3.6 mm long, glabrous; lower glume ca. 1/3 as long as spikelet. Weedy areas, presumably escaping from cultivation; Harris (Brown 24214, SBSC), Bell, Dallas, Travis, and Williamson (Barkworth et al. 2002) cos.; also Galveston Co. (SBSC, TAES); scattered in the s U.S. in CA, FL, GA, LA, OK, and TX. Sep–Nov. Native of Africa. [Panicum maximum Jacq.] This species is an important forage grass, particularly in tropical-subtropical pastures (Gould 1975b). I Urochloa mutica (Forssk.) T.Q. Nguyen, (blunt, pointless), PARA LIVER-SEED GRASS, PARA GRASS. Perennial with culms long, decumbent and rooting at lower nodes, the upright portion usually 90–200(–300) cm tall; inflorescence with 10–30 unbranched, spike-like branches, the branches 2.5–8 cm long; spikelets 2.5–3.5 mm long; lower glume 1/5 –1/3 as long as spikelet. Moist disturbed areas; Jefferson Co. (BRIT, TAES) at se corner of East TX; a Brazos Co. record (Turner et al. 2003) was collected at the Agricultural Experiment Station and is therefore presumably cultivated; mainly s tip of TX; AL, FL, SC, and TX. Sep–Dec. Native of Africa. [Brachiaria mutica (Forssk.) Stapf, Panicum muticum Forssk., Panicum purpurascens Raddi] This species is grown as a forage crop in the tropics (Wipff & Thompson 2003). I Urochloa platyphylla (Munro ex C. Wright) R.D. Webster, (broad-leaved), BROAD-LEAF SIGNAL GRASS, BROAD-LEAF LIVER-SEED GRASS. Coarse annual with decumbent and spreading culm bases, ca. 30–60(–100) cm tall; inflorescence with primary branches unbranched, the branches 3–8 cm long, widely spaced, winged (the plant thus superficially very much resembling Paspalum); spikelets 3.5–4.5(–5) mm long, without a distinct tuft of hairs at base; lower glume 1/4–1/3 as long as spikelet; glumes and lemma of lower floret glabrous. Disturbed areas, roadsides, ditches; in much of the e 1/2 of TX; se U.S. from VA s to FL w to MO, OK, and TX. Apr–Nov. [Brachiaria platyphylla (Munro ex C. Wright) Nash, Paspalum platyphyllum Griseb. not Schult.] See note at U. mosambicensis (following U. texana). Urochloa ramosa (L.) R.D. Webster, (branched), BROWNTOP-MILLET, DIXIE LIVER-SEED GRASS. Annual; culms 25–100 cm tall; inflorescence branches ascending, often rebranched; spikelets (2.4–)
ILLUSTRATED FLORA OF EAST TEXAS 1093 Tridens texanus [HI2, USB] Triplasis purpurea [BB2] Tripogon spicatus [HI1] Tripsacum dactyloides [HI1] Trisetum interruptum [USB] Triticum aestivum [HI1] Urochloa ciliatissima [GO4] Urochloa fusca [GO1]
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KEYS TO THE VASCULAR PLANTS 309 KEY
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KEY TO FERNS AND SIMILAR PLANTS 311
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Key to Families of Gymnosperms KEY
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20. Flowers in a single, small (4-4
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38. Stamens 6 per flower. 39. Peria
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57. Leaves reduced to scales only;
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FERNS AND SIMILAR PLANTS (PTERIDOPH
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ISOETES/ISOETACEAE 323 DIVISION LYC
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Psilotum nudum [KNO] Isoetes butler
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LYCOPODIELLA/LYCOPODIACEAE 327 Beit
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SELAGINELLA/SELAGINELLACEAE 329 Pse
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ILLUSTRATED FLORA OF EAST TEXAS 331
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Palhinhaea cernua [CLU] Selaginella
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leaf blade petiole rhizome root sca
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ASPLENIUM L. SPLEENWORT ASPLENIUM/A
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WOODWARDIA/BLECHNACEAE 339 1. Plant
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MEGASPORE Woodwardia areolata [LUN]
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DRYOPTERIDACEAE 343 AThe family as
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Anemia mexicana Azolla caroliniana
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DRYOPTERIS/DRYOPTERIDACEAE 347 Carr
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WOODSIA/DRYOPTERIDACEAE 351 Tectari
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MARSILEA/MARSILEACEAE 353 etation (
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BOTRYCHIUM/OPHIOGLOSSACEAE 357 REFE
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Nephrolepis exaltata Onoclea sensib
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OPHIOGLOSSUM/OPHIOGLOSSACEAE 361 Op
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PTERIDACEAE 365 and pous or podium,
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Botrychium virginianum Ophioglossum
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CERATOPTERIS/PTERIDACEAE 371 Astrol
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Argyrochosma dealbata [LUN] Astrole
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Cheilanthes feei [LUN] Cheilanthes
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Astrolepis integerrima Astrolepis s
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SALVINIACEAE 379 fern, derived from
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SALVINIA/SALVINIACEAE 381 NM, and N
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Pellaea ovata Pellaea wrightiana IL
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Thelypteris dentata [SM3] STERILE F
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GYMNOSPERMS EPHEDRACEAE 389 AThe te
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DIVISION PINOPHYTA CONIFERS CUPRESS
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JUNIPERUS/CUPRESSACEAE 393 allergen
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Ephedra antisyphilitica [LUN] Junip
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PINUS/PINACEAE 397 clusters at the
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� � � � � � Pinus echin
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Ephedra antisyphilitica Juniperus a
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MAGNOLIOPHYTA 403 consumed in large
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CLASS MONOCOTYLEDONAE MONOCOTYLEDON
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AGAVACEAE 407 produced in North Ame
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HESPERALOE/AGAVACEAE 409 for their
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Acorus calamus [GLE] Hesperaloe eng
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YUCCA/AGAVACEAE 413 snakebite (Verh
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7. Leaves straight or nearly so, la
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Manfreda virginica subsp. lata [HEA
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Manfreda maculosa Manfreda sileri I
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ALISMA/ALISMATACEAE 423 1. Pistils
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Yucca rupicola [LYN] Yucca treculea
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SAGITTARIA/ALISMATACEAE 427 long te
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Echinodorus tenellus [FLN, RHO] Sag
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Yucca treculeana ILLUSTRATED FLORA
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ALLIUM/ALLIACEAE 433 pounds are ant
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ALLIUM/ALLIACEAE 435 Allium ampelop
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STAMENS ILLUSTRATED FLORA OF EAST T
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CAPSULE Allium canadense var. mobil
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var. fraseri-▲ var. hyacinthoides
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AMARYLLIDACEAE 443 and edible (Baye
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COOPERIA/AMARYLLIDACEAE 445 REFEREN
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BULBILS CAPSULE Allium sativum [ANO
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HABRANTHUS/AMARYLLIDACEAE 449 obser
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CAPSULES SEED Habranthus tubispathu
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LYCORIS/AMARYLLIDACEAE 455 usually
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Hymenocallis liriosme [HEA] Leucoju
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Narcissus jonquilla [NIC] Narcissus
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ANTHERICACEAE 461 2000), Anderson,
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ARACEAE 463 hemiepiphytes, and aqua
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ARISAEMA/ARACEAE 465 has been disag
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ORONTIUM/ARACEAE 469 down the insid
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Echeandia flavescens [MCV] Colocasi
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ARECACEAE 473 oped rhizomes present
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ASPARAGACEAE 475 spreading from cul
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BROMELIACEAE 477 jointed pedicels;
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Xanthosoma sagittifolium [BA1] Saba
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APTERIA/BURMANNIACEAE 481 FAMILY RE
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S. minor � S. mexicana-� ILLUST
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UVULARIA/COLCHICACEAE 485 blurred,
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ANTHER CAPSULE PISTIL Apteria aphyl
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CALLISIA/COMMELINACEAE 489 the Haem
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COMMELINA/COMMELINACEAE 491 4. Spat
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SINGLE FLOWER INFLORESCENCE Callisi
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TRADESCANTIA/COMMELINACEAE 495 & Tv
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FRUITING INFLORESCENCE Murdannia nu
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Tradescantia paludosa [AND] Tradesc
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CYPERACEAE 507 become split by grow
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CYPERACEAE 509 10. Scales of spikel
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BULBOSTYLIS/CYPERACEAE 511 1. Spike
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CAREX/CYPERACEAE 517 The following
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CAREX/CYPERACEAE 519 of leaf sheath
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CAREX/CYPERACEAE 521 37. Perigynia
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CAREX/CYPERACEAE 523 58. Achene sho
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CAREX/CYPERACEAE 525 85. Larger per
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Carex abscondita [MAC] Carex albica
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CAREX/CYPERACEAE 531 persists in li
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Carex decomposita [MAC] Carex digit
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Carex triangularis [MAC] Carex tetr
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CYPERUS/CYPERACEAE 559 1974; Baijan
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CYPERUS/CYPERACEAE 561 21. Leaf bla
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CYPERUS/CYPERACEAE 563 fresh; achen
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CYPERUS/CYPERACEAE 565 tively more
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CYPERUS/CYPERACEAE 573 Cyperus invo
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Cyperus esculentus [REE] Cyperus fi
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CYPERUS/CYPERACEAE 577 around the s
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Cyperus iria [GWO] Cyperus lanceola
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Cyperus surinamensis [BT2] Cyperus
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ELEOCHARIS/CYPERACEAE 587 5. Culms
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ELEOCHARIS/CYPERACEAE 589 27. Spike
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ELEOCHARIS/CYPERACEAE 591 43. Scale
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ELEOCHARIS/CYPERACEAE 593 compressa
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ELEOCHARIS/CYPERACEAE 601 ing mats;
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FIMBRISTYLIS Vahl FIMBRY FIMBRISTYL
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Fimbristylis annua [SID] Fimbristyl
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REFERENCES: Coville 1890; Bush 1905
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RHYNCHOSPORA/CYPERACEAE 621 Lipocar
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RHYNCHOSPORA/CYPERACEAE 623 10. Per
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RHYNCHOSPORA/CYPERACEAE 625 34. Cul
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Lipocarpha aristulata [GWO] Lipocar
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SCHOENOPLECTUS/CYPERACEAE 641 exten
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Schoenoplectus pungens [MAS] Schoen
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SCLERIA/CYPERACEAE 649 5. Lowest la
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SCLERIA/CYPERACEAE 651 with a pair
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ERIOCAULON/ERIOCAULACEAE 657 1. Pet
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ERIOCAULON/ERIOCAULACEAE 659 bracts
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� Dioscorea bulbifera [HEB] ILLUS
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HYACINTHACEAE 663 orange zone towar
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Scleria pauciflora ILLUSTRATED FLOR
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� � ILLUSTRATED FLORA OF EAST T
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SCHOENOLIRION/HYACINTHACEAE 669 som
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HYDROCHARITACEAE 671 AA small (116
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HYDRILLIA/HYDROCHARITACEAE 673 marg
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OTTELIA Pers. DUCK-LETTUCE OTTELIA/
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� Vallisneria americana [LAM] Ott
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IRIDACEAE 683 inflorescences with 1
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BELAMCANDA Adans. LEOPARD-LILY, BLA
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IRIS/IRIDACEAE 691 6. Cauline leaf
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Iris brevicaulis [YAT] ILLUSTRATED
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SISYRINCHIUM/IRIDACEAE 697 3. Inflo
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Sisyrinchium ensigerum [CUR] Sisyri
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JUNCUS/JUNCACEAE 703 mainly in wet
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JUNCUS/JUNCACEAE 705 yellowish, lea
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Juncus acuminatus [MAS] Juncus anth
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Juncus dichotomus [GWO] Juncus effu
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Juncus polycephalus [GWO] Juncus re
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var. validus-● var. facinatus-�
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LEMNA/LEMNACEAE 721 Pineywoods and
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SPIRODELA/LEMNACEAE 723 Lemna turio
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NOTE 3 VEINS � TOP VIEW Lemna min
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LILIACEAE 729 The lily family (even
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ERYTHRONIUM/LILIACEAE 731 17. Tepal
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Wolffia columbiana [GWO, MIB] Wolff
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LIMNOCHARITACEAE 735 1. Perianth wh
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THALIA/MARANTACEAE 737 pollen prese
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MELANTHIACEAE 739 AA small family (
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Lilium philippinense [GCH] SEED FOL
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VERATRUM/MELANTHIACEAE 745 circumsc
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Zigadenus glaberrimus [GLE] Schoeno
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ALETRIS/NARTHECIACEAE 749 Tofieldia
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DASYLIRION Zucc. SOTOL, SHAGGY-LILY
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FLOWER Aletris aurea [CO1] Aletris
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ORCHIDACEAE 755 45 occur in East TX
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CALOPOGON/ORCHIDACEAE 757 5-25 flow
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CLEISTES/ORCHIDACEAE 759 long; lip
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CYPRIPEDIUM/ORCHIDACEAE 761 while t
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Stenanthium gramineum Toxicoscordio
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Calopogon tuberosus [FNA] Corallorh
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HEXALECTRIS/ORCHIDACEAE 767 rain. A
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Habenaria quinqueseta [LUN] Habenar
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PLATANTHERA/ORCHIDACEAE 771 PLATANT
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PLATANTHERA/ORCHIDACEAE 773 Platant
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SPIRANTHES/ORCHIDACEAE 779 mm wide,
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SPIRANTHES/ORCHIDACEAE 781 12. Lip
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Platanthera lacera [YAT] Pogonia op
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SPIRANTHES/ORCHIDACEAE 789 narrowes
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TRIPHORA/ORCHIDACEAE 791 an example
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LEAF collar rhizome blade ligule sh
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POACEAE 797 Dissection is usually n
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POACEAE 799 rower and often differe
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POACEAE 801 6. Lemmas awned, the tw
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POACEAE 803 30. Lemma veins none to
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POACEAE 805 13. Inflorescences narr
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POACEAE 807 12. Lemma awnless; infl
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POACEAE 809 inflorescence solitary,
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POACEAE 811 29. Spikelets loosely a
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POACEAE 813 20. Basal leaves usuall
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AEGILOPS/POACEAE 815 23. Spikelets
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AGROSTIS/POACEAE 817 Agrostis hyema
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ANDROPOGON/POACEAE 821 colonizing e
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ANTHENANTIA/POACEAE 825 to be the e
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ARISTIDA/POACEAE 829 9. Lemma thick
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ARISTIDA/POACEAE 831 BRIT), Newton
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Anthoxanthum aristatum [BB2] Anthox
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AXONOPUS/POACEAE 839 duced to the U
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Avena fatua [HI1] Arundo donax [LYN
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BOTHRIOCHLOA/POACEAE 843 var. barbi
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Aristida purpurea (all vars.) ILLUS
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Axonopus compressus [HI1] Axonopus
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BOUTELOUA/POACEAE 849 state grass b
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Bothriochloa ischaemum var. songari
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Bouteloua pectinata [BOT, HEA] Bout
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BROMUS/POACEAE 857 8. Lemmas and gl
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BUCHLOE/POACEAE 861 ticularly in th
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CHASMANTHIUM/POACEAE 865 [C. caroli
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CHLORIS/POACEAE 871 East TX; mainly
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Chasmanthium sessiliflorum [USB] Ch
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CORTADERIA/POACEAE 875 of the inflo
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DANTHONIA/POACEAE 881 Dactylocteniu
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Cynodon dactylon [REE] Cynosurus ec
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DICHANTHELIUM/POACEAE 885 AA genus
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DICHANTHELIUM/POACEAE 887 20. Inter
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DICHANTHELIUM/POACEAE 889 26. Leaf
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Danthonia spicata [USB] Desmazeria
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DICHANTHELIUM/POACEAE 897 and Harri
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Dichanthelium acuminatum (all vars.
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DICHANTHELIUM/POACEAE 905 that this
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DIGITARIA/POACEAE 909 2. Main axis
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Dichanthelium sphaerocarpon [HI] IL
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DIGITARIA/POACEAE 913 with papilla-
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Digitaria bicornis [HEB] ILLUSTRATE
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Digitaria insularis [HI1] Digitaria
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EHRHARTA/POACEAE 921 spikelets (2.5
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ELIONURUS/POACEAE 923 Since this co
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Echinochloa crus-gali [REE] Echinoc
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ENTEROPOGON Nees UMBRELLA GRASS ENT
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ERAGROSTIS/POACEAE 931 8. Infloresc
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ERAGROSTIS/POACEAE 933 12. Pedicels
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Elymus repens [HI1] Elymus smithii
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Eragrostis japonica [USB] ILLUSTRAT
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ERIONEURON/POACEAE 945 Shaw and Sme
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� � � ILLUSTRATED FLORA OF EA
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FESTUCA/POACEAE 951 rachilla glabro
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GYMNOPOGON/POACEAE 955 Glyceria sep
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HILARIA/POACEAE 957 REFERENCES: Eme
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ILLUSTRATED FLORA OF EAST TEXAS Fes
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KOELERIA/POACEAE 963 Andropogoneae,
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Heteropogon melanocarpus [HI1] Hord
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LEPTOCHLOA/POACEAE 967 Leersia oryz
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ILLUSTRATED FLORA OF EAST TEXAS Koe
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ILLUSTRATED FLORA OF EAST TEXAS Hor
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LOLIUM/POACEAE 973 Charmet et al. 1
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Leptochloa nealleyi [USB] Lolium ri
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MICROSTEGIUM/POACEAE 977 AA variabl
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ILLUSTRATED FLORA OF EAST TEXAS Lep
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Lolium temulentum [USB] Melica muti
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MUHLENBERGIA/POACEAE 983 10. Lemma
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MUHLENBERGIA/POACEAE 985 and Prairi
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ILLUSTRATED FLORA OF EAST TEXAS Mon
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ILLUSTRATED FLORA OF EAST TEXAS Mic
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 993 fact that gener
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PANICUM/POACEAE 995 17. Spikelets 2
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Oryza sativa [HI1] Panicum anceps [
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 1001 Panicum hallii
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Panicum trichoides [HI2] ILLUSTRATE
Page 701 and 702:
PASPALUM/POACEAE 1009 5. Spikelets
Page 703 and 704:
PASPALUM/POACEAE 1011 27. Axils of
Page 705 and 706:
Panicum virgatum [RCA] Paspalum acu
Page 707 and 708:
Page 709 and 710:
Page 711 and 712:
PASPALUM/POACEAE 1019 loam soils; w
Page 713 and 714:
Page 715 and 716:
Page 717 and 718:
PENNISETUM/POACEAE 1025 unit as in
Page 719 and 720:
Paspalum repens [USB] ILLUSTRATED F
Page 721 and 722:
PHALARIS/POACEAE 1029 landscapes (B
Page 723 and 724:
Paspalum pubiflorum (both vars.) IL
Page 725 and 726:
Pennisetum purpureum [HUT] Penniset
Page 727 and 728:
POA/POACEAE 1035 around awn base; l
Page 729 and 730:
POA/POACEAE 1037 long. Forests, flo
Page 731 and 732:
Page 733 and 734: ILLUSTRATED FLORA OF EAST TEXAS 104
Page 737 and 738: SACCHARUM/POACEAE 1045 1. Awn of le
Page 739 and 740: SCHEDONNARDUS/POACEAE 1047 lower gl
Page 745 and 746: SETARIA/POACEAE 1053 Paspalidium (W
Page 747 and 748: SETARIA/POACEAE 1055 14. Leaf blade
Page 749 and 750: Schizachyrium tenerum [HI1, USB] Sc
Page 755 and 756: SORGHUM/POACEAE 1063 2000) indicate
Page 757 and 758: SPARTINA/POACEAE 1065 Americas it [
Page 759 and 760: Sorghastrum elliottii [HI1] Sorghas
Page 763 and 764: SPOROBOLUS/POACEAE 1071 7. Lemma ap
Page 767 and 768: STEINCHISMA/POACEAE 1075 Sporobolus
Page 769 and 770: Sphenopholis obtusata (both vars.)
Page 771 and 772: Sporobolus neglectus [HI1] Sporobol
Page 773 and 774: TRICHONEURA/POACEAE 1081 guish it f
Page 775 and 776: Stenotaphrum secundatum [USB] Trich
Page 781 and 782: TRISETUM/POACEAE 1089 low areas; ne
Page 783: UROCHLOA/POACEAE 1091 AA C4 genus o
Page 787 and 788: Tridens eragrostoides Tridens flavu
Page 789 and 790: VULPIA/POACEAE 1097 2. Lemmas (exce
Page 795 and 796: ZIZANIOPSIS/POACEAE 1103 (1999) (po
Page 799 and 800: EICHHORNIA/PONTEDERIACEAE 1107 REFE
Page 801 and 802: Zoysia japonica [HOL] Zoysia pacifi
Page 803 and 804: POTAMOGETON/POTAMOGETONACEAE 1111 1
Page 805 and 806: POTAMOGETON/POTAMOGETONACEAE 1113 P
Page 811 and 812: SMILAX/SMILACACEAE 1119 times woody
Page 813 and 814: SMILAX/SMILACACEAE 1121 30(-60) mm
Page 815 and 816: Stuckenia pectinata [MAS] ILLUSTRAT
Page 819 and 820: THEMIDACEAE 1127 cover for various
Page 821 and 822: Smilax rotundifolia [JEM] ILLUSTRAT
Page 823 and 824: TRILLIUM/TRILLIACEAE 1131 Schilling
Page 825 and 826: TRILLIUM/TRILLIACEAE 1133 taxonomic
Page 827 and 828: Triantha racemosa [GLE] Trillium lu
Page 829 and 830: TYPHA/TYPHACEAE 1137 REFERENCES: Wi
Page 831 and 832: XYRIS/XYRIDACEAE 1139 cone-like spi
Page 833 and 834: XYRIS/XYRIDACEAE 1141 11. Plant bas
Page 835 and 836:
Sparganium americanum Typha latifol
Page 837 and 838:
Typha domingensis[ BT3] Typha latif
Page 839 and 840:
Xyris drummondii [SID] Xyris fimbri
Page 841 and 842:
Xyris smalliana [SID] ILLUSTRATED F
Page 843:
Xyris caroliniana Xyris difformis (
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