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keys to the vascular plants of east texas - Botanical Research ...

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658 ERIOCAULACEAE/ERIOCAULON<br />

basal rosette, linear, sheath-like at base; inflorescence a compact, hemispherical <strong>to</strong> globose head<br />

terminating an elongate scape; bracts and perianth parts <strong>of</strong>ten with conspicuous, clavate, whitish<br />

hairs; flowers small (ca. 3 mm or less long) and crowded <strong>to</strong>ge<strong>the</strong>r; sepals 2; petals 2; male<br />

flowers with (3–)4(–6) stamens; carpels 2.<br />

AA genus <strong>of</strong> ca. 400 species <strong>of</strong> tropical and warm areas, including Japan and North America,<br />

most aquatic or in wet, acidic habitats (Kral 2000b). The dried inflorescences are sometimes<br />

used in floral arrangements (Judd et al. 1999). (Greek: erion, wool, and caulos, a stalk, from <strong>the</strong><br />

wool at <strong>the</strong> base <strong>of</strong> <strong>the</strong> scape in <strong>the</strong> first named species)<br />

REFERENCES: Kral 1966b; Watson et al. 1994.<br />

1. Heads 3–4 mm broad at flowering time, dark gray or gray-green except for pale “rims” <strong>of</strong> white<br />

ciliate perianth parts and receptacular bracts and pale outer involucral bracts; <strong>plants</strong> diminutive,<br />

<strong>the</strong> mature scapes only 5–8 cm long; receptacle surface glabrous; species rare in East TX<br />

_________________________________________________________________________ E. koernickianum<br />

1. Heads 5–20 mm broad at flowering time, white or gray; <strong>plants</strong> usually larger, <strong>the</strong> mature scapes<br />

5–110 cm long; receptacle surface hairy; including species widespread and common (in appropriate<br />

habitats) in East TX.<br />

2. Heads s<strong>of</strong>t, much compressed when pressed and dried; sheaths <strong>of</strong> <strong>the</strong> scapes exceeding all<br />

or most leaves in length; involucral bracts grayish, obtuse or rounded apically; receptacular<br />

bracts grayish, acute apically; petals <strong>of</strong> pistillate flowers with inner (= adaxial) surface villous;<br />

clavate hairs <strong>of</strong> perianth with all cells whitened and opaque; <strong>plants</strong> <strong>of</strong> aquatic or wet situations,<br />

flowering and fruiting in winter, spring, or early summer.<br />

3. Heads 10–20 mm broad at flowering time; scapes 20–70 cm long; leaves 5–30 cm long;<br />

roots usually 1 mm or more broad; corollas <strong>of</strong> staminate flowers with lobes conspicuously<br />

unequal ________________________________________________________________ E. compressum<br />

3. Heads usually ca. 5–10 mm broad at flowering time; scapes usually 5–30 cm long; leaves 1–<br />

5(–7) cm long; roots usually 1 mm or less broad; corollas <strong>of</strong> staminate flowers with lobes<br />

subequal ____________________________________________________________________ E. texense<br />

2. Heads very hard, only slightly compressed when pressed and dried; sheaths <strong>of</strong> <strong>the</strong> scapes<br />

usually exceeded in length by most <strong>of</strong> <strong>the</strong> leaves; involucral bracts straw-colored, acute apically;<br />

receptacular bracts pale, narrowly acute <strong>to</strong> acuminate apically; petals <strong>of</strong> pistillate flowers with<br />

inner surface glabrous; clavate hairs <strong>of</strong> <strong>the</strong> perianth with terminal cells whitened and opaque,<br />

but some or all <strong>of</strong> such hairs with basal cells not white but transparent; <strong>plants</strong> <strong>of</strong> moist but<br />

seldom aquatic or permanently wet situations, flowering late spring or summer and fruiting<br />

in summer and fall ___________________________________________________________ E. decangulare<br />

Eriocaulon compressum Lam., (compressed, flattened), FLATTENED PIPEWORT. Leaves 5–30 cm<br />

long; mature scapes 20–70 cm long; mature (flowering) heads hemispherical or globose, 10–20<br />

mm broad, chalk white except for <strong>the</strong> dark exserted tips <strong>of</strong> <strong>the</strong> receptacular bracts and an<strong>the</strong>rs;<br />

outer involucral bracts 2–3 mm long, obtuse or rounded, grayish-translucent; surface <strong>of</strong> receptacle<br />

with multicellular, translucent hairs; receptacular bracts acute; sepals translucent, with<br />

clavate white hairs on <strong>the</strong> back. Sands or sandy peats <strong>of</strong> wet areas such as acid ponds, swamps,<br />

and pine savannahs; New<strong>to</strong>n (BRIT), Hardin (TAES), Anderson (TAMU), Jasper, Tyler (VDB),<br />

and Jefferson (Turner et al. 2003) cos. in <strong>the</strong> Pineywoods; also n Gulf Prairies and Marshes; se<br />

U.S. from VA s <strong>to</strong> FL w <strong>to</strong> TX. Spring. Kral (1966b) indicated that “This is perhaps <strong>the</strong> showiest<br />

<strong>of</strong> all <strong>the</strong> Eriocaulaceae <strong>of</strong> <strong>the</strong> sou<strong>the</strong>astern United States, in springtime so abundantly decorating<br />

<strong>the</strong> shallow water <strong>of</strong> pinelands as <strong>to</strong> appear like a shower <strong>of</strong> white confetti.” This is <strong>the</strong> most<br />

aquatic <strong>of</strong> <strong>the</strong> species occurring in East TX (Kral 2000b). (TOES 1993: V) �<br />

Eriocaulon decangulare L., (ten-angled), TEN-ANGLE PIPEWORT, PIPEWORT. Leaves 10–40 cm long;<br />

mature scapes 30–110 cm long; mature heads subglobose, 10–20 mm broad, dull white, hard;<br />

lowermost flowers and receptacular bracts reflexed and obscuring <strong>the</strong> subtending involucral

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