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keys to the vascular plants of east texas - Botanical Research ...

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312 KEY TO FERNS AND SIMILAR PLANTS<br />

18. Plants s<strong>to</strong>ut, <strong>to</strong> 1 m or more tall; leaf blades triangular in<br />

shape or nearly so, with 3 main divisions (each division<br />

usually bipinnate); petioles greenish or ± straw-colored;<br />

stems and petiole bases with slender hairs but without<br />

scales ______________________ (Pteridium) Dennstaedtiaceae<br />

18. Plants slender, usually 0.1–0.5 m tall; leaf blades nei<strong>the</strong>r<br />

triangular nor with 3 main divisions; petioles <strong>of</strong>ten dark<br />

brown or black; stems and petiole bases generally with<br />

scales ________________________________________ Pteridaceae<br />

17. Leaf margins never recurved <strong>to</strong> form a false indusium; sori<br />

variously located on <strong>the</strong> abaxial (= beneath) leaf surfaces,<br />

<strong>of</strong>ten near veins, occasionally near <strong>the</strong> margins <strong>of</strong> <strong>the</strong> leaves;<br />

adaxial leaf surfaces without stellate or coarsely ciliate<br />

scales.<br />

19. Sori elongate along <strong>the</strong> veins; leaf blades 1-pinnate _____ (Asplenium)<br />

Aspleniaceae<br />

19. Sori round or variously shaped, not elongate (in our species,<br />

except elongate in Athyrium with leaf blades 2pinnate-pinnatifid);<br />

leaf blades in our species 1-pinnatifid,<br />

1-pinnate, or more than 1-pinnate.<br />

20. Leaf blades in our species only 1-pinnate or 1-pinnatifid,<br />

<strong>the</strong> pinnae not fur<strong>the</strong>r subdivided in<strong>to</strong><br />

pinnules.<br />

21. Leaf blades only pinnatifid (in our species), divided<br />

nearly but not all <strong>the</strong> way <strong>to</strong> <strong>the</strong> rachis<br />

(= midrib); pinnae essentially as wide at base<br />

as <strong>to</strong>wards <strong>the</strong>ir tips; margins <strong>of</strong> pinnae (in our<br />

species) without any teeth or basal auricle; indusia<br />

lacking _________________ (Pleopeltis) Polypodiaceae<br />

21. Leaf blades pinnate, divided all <strong>the</strong> way <strong>to</strong> <strong>the</strong><br />

rachis; pinnae narrowed basally <strong>to</strong> a very narrow<br />

petiole-like attachment <strong>to</strong> <strong>the</strong> rachis; pinnae<br />

with small teeth on <strong>the</strong> margins, sometimes<br />

with basal auricles; indusia present (but sometimes<br />

shriveled at maturity) ____________ Dryopteridaceae<br />

20. Leaf blades more than 1-pinnate, <strong>the</strong> pinnae <strong>the</strong>mselves<br />

ei<strong>the</strong>r pinnate or pinnatifid.<br />

22. Lower leaf surfaces with at l<strong>east</strong> some transparent<br />

needle-like hairs (particularly on <strong>the</strong><br />

costae and veins, sometimes even on <strong>the</strong> intervening<br />

tissue or on <strong>the</strong> indusia—use hand<br />

lens or scope); ultimate leaf segments <strong>of</strong>ten<br />

(but not always) entire; petioles with 2 crescent-shaped<br />

<strong>vascular</strong> bundles basally, distally<br />

<strong>the</strong>se united in<strong>to</strong> a U-shape ___________ Thelypteridaceae<br />

22. Lower leaf surfaces without transparent<br />

needle-like hairs; ultimate leaf segments variously<br />

incised, serrate, dentate, crenate, or lobulate,<br />

not entire; petioles with 2 or more round<br />

or oblong <strong>vascular</strong> bundles ____________ Dryopteridaceae

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