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500 CONVALLARIACEAE key. While recognizing it as a distinct species, Faden (2000b) indicated that this is “… clearly Anderson and Woodson’s weakest species, and D.T. MacRoberts (1979) may be correct in treating it as a variety of Tradescantia ohiensis.” This species can be cultivated year-round in greenhouses, since the plants apparently do not require a winter dormancy (Faden 2000b). Tradescantia reverchonii Bush, (for Julien Reverchon, 1837–1905, a French-American immigrant to Dallas and important botanical collector of early TX), REVERCHON’S SPIDERWORT, GRASS-VIO- LET. Roots thick, fleshy, conspicuously felty with red-brown hairs easily visible to the naked eye; stems 30–105 cm tall, erect or ascending, simple or infrequently branched, usually rather densely pilose with somewhat matted or tangled hairs; sepals with eglandular or both eglandular and glandular hairs; petals bright blue (rarely rose or white). Sandy soils, open woods; widespread in the s and e parts of East TX, also collected at Dallas by Reverchon (Mahler 1988) but not found there since; also Gulf Prairies and Marshes, South TX Plains, and e Edwards Plateau; AR, LA, and TX. Mar–Jul. Tradescantia subacaulis Bush, (almost without a stem), STEMLESS SPIDERWORT. Roots tuberousthickened; stems spreading, 10–35 cm long, densely matted-pilose throughout; sepals with glandular and eglandular hairs; petals usually bright blue, occasionally pink. Loose sandy soils, open woods or open ground; widespread in e 1/2 of TX; endemic to TX (Faden 2000b; Carr 2002b, 2002c). Late Mar–Jun. E Tradescantia tharpii E.S. Anderson & Woodson, (for Benjamin Carroll Tharp, 1885–1964, botanist at Univ. of TX and author of Structure of Texas Vegetation East of the 98th Meridian), THARP’S SPIDERWORT. Plant long-pilose throughout, usually densely so; stems 2–7 cm tall in flower, to 20(–30) cm in fruit; stems rarely branching, often initially acaulescent; sepals with only eglandular pubescence; petals deep rose or purple, sometimes blue. Sandy clay or rarely silty clay soils, rocky prairies, open woods, or open ground; Collin, Denton, and Dallas (BRIT) cos. in the Blackland Prairie; while we have seen specimens from only a few counties, Hatch et al. (1990) cited vegetational areas 4, 5, 7, and 8, and the range map in Faden (2000b) shows the species occurring throughout East TX; KS, MO, OK, and TX. Late Mar–Apr. CONVALLARIACEAE Horan. LILY-OF-THE-VALLEY FAMILY Rhizomatous perennial herbs, acaulescent or with arching to erect-arching stems; leaves all basal or cauline; inflorescences terminal or axillary; flowers perfect, 3-merous; perianth of distinct or fused segments; stamens 6; ovary superior or partly inferior; fruit a berry. AA small family (17 genera and 130 species) found in North and Central America, Europe, and Asia (Conran & Tamura 1998). The family placement of the genera treated here as Convallariaceae has been extremely variable. Many authorities in the past have put them in a broadly defined and clearly polyphyletic (but practical) Liliaceae (e.g., Correll & Johnston 1970; Cronquist 1988; Diggs et al. 1999) based on superficial similarities in flower structure to that of the genus Lilium. However, molecular analyses (Chase et al. 1995a, 2000; Fay et al. 2000) leave little doubt that the group belongs in order Asparagales. Judd et al. (1999) and Yamashita and Tamura (2000) placed the genera in the Convallariaceae, while Rudall et al. (2000a), Judd et al. (2002), and Judd (2003) treated them in a broadly interpreted Ruscaceae. More recently, the Angiosperm Phylogeny Group (APG II 2003) suggested placing them either in Ruscaceae or in a very broadly defined Asparagaceae (along with such families as Agavaceae). While placement in Asparagales now seems clear, there has been disagreement in molecular analyses of the complex that could possibly affect family circumscription (e.g., Rudall et al. 2000a; Yamashita & Tamura 2000). Until taxonomy of the group is more settled, we are following Conran and Tamura (1998) and Yamashita and Tamura (2000) in recognizing the Convallariaceae at the
ILLUSTRATED FLORA OF EAST TEXAS 501 Tradescantia hirsutiflora [HEA] Tradescantia humilis [HEA] SEPALS Tradescantia ohiensis [BB2] Tradescantia leiandra [HEB] Tradescantia occidentalis [FNA]
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KEYS TO THE VASCULAR PLANTS 309 KEY
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KEY TO FERNS AND SIMILAR PLANTS 311
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Key to Families of Gymnosperms KEY
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20. Flowers in a single, small (4-4
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38. Stamens 6 per flower. 39. Peria
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57. Leaves reduced to scales only;
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FERNS AND SIMILAR PLANTS (PTERIDOPH
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ISOETES/ISOETACEAE 323 DIVISION LYC
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Psilotum nudum [KNO] Isoetes butler
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LYCOPODIELLA/LYCOPODIACEAE 327 Beit
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SELAGINELLA/SELAGINELLACEAE 329 Pse
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ILLUSTRATED FLORA OF EAST TEXAS 331
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Palhinhaea cernua [CLU] Selaginella
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leaf blade petiole rhizome root sca
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ASPLENIUM L. SPLEENWORT ASPLENIUM/A
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WOODWARDIA/BLECHNACEAE 339 1. Plant
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MEGASPORE Woodwardia areolata [LUN]
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DRYOPTERIDACEAE 343 AThe family as
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Anemia mexicana Azolla caroliniana
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DRYOPTERIS/DRYOPTERIDACEAE 347 Carr
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WOODSIA/DRYOPTERIDACEAE 351 Tectari
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MARSILEA/MARSILEACEAE 353 etation (
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BOTRYCHIUM/OPHIOGLOSSACEAE 357 REFE
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Nephrolepis exaltata Onoclea sensib
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OPHIOGLOSSUM/OPHIOGLOSSACEAE 361 Op
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PTERIDACEAE 365 and pous or podium,
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Botrychium virginianum Ophioglossum
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CERATOPTERIS/PTERIDACEAE 371 Astrol
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Argyrochosma dealbata [LUN] Astrole
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Cheilanthes feei [LUN] Cheilanthes
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Astrolepis integerrima Astrolepis s
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SALVINIACEAE 379 fern, derived from
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SALVINIA/SALVINIACEAE 381 NM, and N
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Pellaea ovata Pellaea wrightiana IL
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Thelypteris dentata [SM3] STERILE F
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GYMNOSPERMS EPHEDRACEAE 389 AThe te
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DIVISION PINOPHYTA CONIFERS CUPRESS
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JUNIPERUS/CUPRESSACEAE 393 allergen
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Ephedra antisyphilitica [LUN] Junip
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PINUS/PINACEAE 397 clusters at the
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� � � � � � Pinus echin
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Ephedra antisyphilitica Juniperus a
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MAGNOLIOPHYTA 403 consumed in large
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CLASS MONOCOTYLEDONAE MONOCOTYLEDON
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AGAVACEAE 407 produced in North Ame
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HESPERALOE/AGAVACEAE 409 for their
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Acorus calamus [GLE] Hesperaloe eng
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YUCCA/AGAVACEAE 413 snakebite (Verh
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7. Leaves straight or nearly so, la
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Manfreda virginica subsp. lata [HEA
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Manfreda maculosa Manfreda sileri I
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ALISMA/ALISMATACEAE 423 1. Pistils
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Yucca rupicola [LYN] Yucca treculea
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SAGITTARIA/ALISMATACEAE 427 long te
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Echinodorus tenellus [FLN, RHO] Sag
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Yucca treculeana ILLUSTRATED FLORA
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ALLIUM/ALLIACEAE 433 pounds are ant
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ALLIUM/ALLIACEAE 435 Allium ampelop
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STAMENS ILLUSTRATED FLORA OF EAST T
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CAPSULE Allium canadense var. mobil
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var. fraseri-▲ var. hyacinthoides
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AMARYLLIDACEAE 443 and edible (Baye
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COOPERIA/AMARYLLIDACEAE 445 REFEREN
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BULBILS CAPSULE Allium sativum [ANO
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Page 143 and 144: CAPSULES SEED Habranthus tubispathu
Page 145 and 146: ILLUSTRATED FLORA OF EAST TEXAS 453
Page 147 and 148: LYCORIS/AMARYLLIDACEAE 455 usually
Page 149 and 150: Hymenocallis liriosme [HEA] Leucoju
Page 151 and 152: Narcissus jonquilla [NIC] Narcissus
Page 153 and 154: ANTHERICACEAE 461 2000), Anderson,
Page 155 and 156: ARACEAE 463 hemiepiphytes, and aqua
Page 157 and 158: ARISAEMA/ARACEAE 465 has been disag
Page 161 and 162: ORONTIUM/ARACEAE 469 down the insid
Page 163 and 164: Echeandia flavescens [MCV] Colocasi
Page 165 and 166: ARECACEAE 473 oped rhizomes present
Page 167 and 168: ASPARAGACEAE 475 spreading from cul
Page 169 and 170: BROMELIACEAE 477 jointed pedicels;
Page 171 and 172: Xanthosoma sagittifolium [BA1] Saba
Page 173 and 174: APTERIA/BURMANNIACEAE 481 FAMILY RE
Page 175 and 176: S. minor � S. mexicana-� ILLUST
Page 177 and 178: UVULARIA/COLCHICACEAE 485 blurred,
Page 179 and 180: ANTHER CAPSULE PISTIL Apteria aphyl
Page 181 and 182: CALLISIA/COMMELINACEAE 489 the Haem
Page 183 and 184: COMMELINA/COMMELINACEAE 491 4. Spat
Page 185 and 186: SINGLE FLOWER INFLORESCENCE Callisi
Page 187 and 188: TRADESCANTIA/COMMELINACEAE 495 & Tv
Page 189 and 190: FRUITING INFLORESCENCE Murdannia nu
Page 191: ILLUSTRATED FLORA OF EAST TEXAS 499
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Page 199 and 200: CYPERACEAE 507 become split by grow
Page 201 and 202: CYPERACEAE 509 10. Scales of spikel
Page 203 and 204: BULBOSTYLIS/CYPERACEAE 511 1. Spike
Page 209 and 210: CAREX/CYPERACEAE 517 The following
Page 211 and 212: CAREX/CYPERACEAE 519 of leaf sheath
Page 213 and 214: CAREX/CYPERACEAE 521 37. Perigynia
Page 215 and 216: CAREX/CYPERACEAE 523 58. Achene sho
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Page 219 and 220: Carex abscondita [MAC] Carex albica
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Carex triangularis [MAC] Carex tetr
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CYPERUS/CYPERACEAE 559 1974; Baijan
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CYPERUS/CYPERACEAE 561 21. Leaf bla
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CYPERUS/CYPERACEAE 563 fresh; achen
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CYPERUS/CYPERACEAE 565 tively more
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CYPERUS/CYPERACEAE 573 Cyperus invo
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Cyperus esculentus [REE] Cyperus fi
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CYPERUS/CYPERACEAE 577 around the s
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Cyperus iria [GWO] Cyperus lanceola
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Cyperus surinamensis [BT2] Cyperus
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ELEOCHARIS/CYPERACEAE 587 5. Culms
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ELEOCHARIS/CYPERACEAE 589 27. Spike
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ELEOCHARIS/CYPERACEAE 591 43. Scale
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ELEOCHARIS/CYPERACEAE 593 compressa
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ELEOCHARIS/CYPERACEAE 601 ing mats;
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FIMBRISTYLIS Vahl FIMBRY FIMBRISTYL
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Fimbristylis annua [SID] Fimbristyl
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REFERENCES: Coville 1890; Bush 1905
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RHYNCHOSPORA/CYPERACEAE 621 Lipocar
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RHYNCHOSPORA/CYPERACEAE 623 10. Per
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RHYNCHOSPORA/CYPERACEAE 625 34. Cul
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Lipocarpha aristulata [GWO] Lipocar
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SCHOENOPLECTUS/CYPERACEAE 641 exten
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Schoenoplectus pungens [MAS] Schoen
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SCLERIA/CYPERACEAE 649 5. Lowest la
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SCLERIA/CYPERACEAE 651 with a pair
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ERIOCAULON/ERIOCAULACEAE 657 1. Pet
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ERIOCAULON/ERIOCAULACEAE 659 bracts
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� Dioscorea bulbifera [HEB] ILLUS
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HYACINTHACEAE 663 orange zone towar
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Scleria pauciflora ILLUSTRATED FLOR
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� � ILLUSTRATED FLORA OF EAST T
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SCHOENOLIRION/HYACINTHACEAE 669 som
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HYDROCHARITACEAE 671 AA small (116
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HYDRILLIA/HYDROCHARITACEAE 673 marg
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OTTELIA Pers. DUCK-LETTUCE OTTELIA/
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� Vallisneria americana [LAM] Ott
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IRIDACEAE 683 inflorescences with 1
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BELAMCANDA Adans. LEOPARD-LILY, BLA
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IRIS/IRIDACEAE 691 6. Cauline leaf
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Iris brevicaulis [YAT] ILLUSTRATED
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SISYRINCHIUM/IRIDACEAE 697 3. Inflo
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Sisyrinchium ensigerum [CUR] Sisyri
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JUNCUS/JUNCACEAE 703 mainly in wet
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JUNCUS/JUNCACEAE 705 yellowish, lea
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Juncus acuminatus [MAS] Juncus anth
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Juncus dichotomus [GWO] Juncus effu
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Juncus polycephalus [GWO] Juncus re
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var. validus-● var. facinatus-�
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LEMNA/LEMNACEAE 721 Pineywoods and
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SPIRODELA/LEMNACEAE 723 Lemna turio
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NOTE 3 VEINS � TOP VIEW Lemna min
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LILIACEAE 729 The lily family (even
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ERYTHRONIUM/LILIACEAE 731 17. Tepal
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Wolffia columbiana [GWO, MIB] Wolff
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LIMNOCHARITACEAE 735 1. Perianth wh
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THALIA/MARANTACEAE 737 pollen prese
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MELANTHIACEAE 739 AA small family (
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Lilium philippinense [GCH] SEED FOL
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VERATRUM/MELANTHIACEAE 745 circumsc
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Zigadenus glaberrimus [GLE] Schoeno
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ALETRIS/NARTHECIACEAE 749 Tofieldia
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DASYLIRION Zucc. SOTOL, SHAGGY-LILY
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FLOWER Aletris aurea [CO1] Aletris
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ORCHIDACEAE 755 45 occur in East TX
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CALOPOGON/ORCHIDACEAE 757 5-25 flow
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CLEISTES/ORCHIDACEAE 759 long; lip
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CYPRIPEDIUM/ORCHIDACEAE 761 while t
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Stenanthium gramineum Toxicoscordio
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Calopogon tuberosus [FNA] Corallorh
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HEXALECTRIS/ORCHIDACEAE 767 rain. A
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Habenaria quinqueseta [LUN] Habenar
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PLATANTHERA/ORCHIDACEAE 771 PLATANT
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PLATANTHERA/ORCHIDACEAE 773 Platant
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SPIRANTHES/ORCHIDACEAE 779 mm wide,
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SPIRANTHES/ORCHIDACEAE 781 12. Lip
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Platanthera lacera [YAT] Pogonia op
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SPIRANTHES/ORCHIDACEAE 789 narrowes
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TRIPHORA/ORCHIDACEAE 791 an example
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LEAF collar rhizome blade ligule sh
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POACEAE 797 Dissection is usually n
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POACEAE 799 rower and often differe
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POACEAE 801 6. Lemmas awned, the tw
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POACEAE 803 30. Lemma veins none to
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POACEAE 805 13. Inflorescences narr
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POACEAE 807 12. Lemma awnless; infl
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POACEAE 809 inflorescence solitary,
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POACEAE 811 29. Spikelets loosely a
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POACEAE 813 20. Basal leaves usuall
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AEGILOPS/POACEAE 815 23. Spikelets
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AGROSTIS/POACEAE 817 Agrostis hyema
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ANDROPOGON/POACEAE 821 colonizing e
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ANTHENANTIA/POACEAE 825 to be the e
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ARISTIDA/POACEAE 829 9. Lemma thick
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ARISTIDA/POACEAE 831 BRIT), Newton
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Anthoxanthum aristatum [BB2] Anthox
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AXONOPUS/POACEAE 839 duced to the U
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Avena fatua [HI1] Arundo donax [LYN
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BOTHRIOCHLOA/POACEAE 843 var. barbi
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Aristida purpurea (all vars.) ILLUS
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Axonopus compressus [HI1] Axonopus
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BOUTELOUA/POACEAE 849 state grass b
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Bothriochloa ischaemum var. songari
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Bouteloua pectinata [BOT, HEA] Bout
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BROMUS/POACEAE 857 8. Lemmas and gl
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BUCHLOE/POACEAE 861 ticularly in th
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CHASMANTHIUM/POACEAE 865 [C. caroli
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CHLORIS/POACEAE 871 East TX; mainly
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Chasmanthium sessiliflorum [USB] Ch
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CORTADERIA/POACEAE 875 of the inflo
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DANTHONIA/POACEAE 881 Dactylocteniu
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Cynodon dactylon [REE] Cynosurus ec
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DICHANTHELIUM/POACEAE 885 AA genus
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DICHANTHELIUM/POACEAE 887 20. Inter
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DICHANTHELIUM/POACEAE 889 26. Leaf
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Danthonia spicata [USB] Desmazeria
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DICHANTHELIUM/POACEAE 897 and Harri
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Dichanthelium acuminatum (all vars.
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DICHANTHELIUM/POACEAE 905 that this
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DIGITARIA/POACEAE 909 2. Main axis
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Dichanthelium sphaerocarpon [HI] IL
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DIGITARIA/POACEAE 913 with papilla-
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Digitaria bicornis [HEB] ILLUSTRATE
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Digitaria insularis [HI1] Digitaria
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EHRHARTA/POACEAE 921 spikelets (2.5
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ELIONURUS/POACEAE 923 Since this co
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Echinochloa crus-gali [REE] Echinoc
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ENTEROPOGON Nees UMBRELLA GRASS ENT
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ERAGROSTIS/POACEAE 931 8. Infloresc
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ERAGROSTIS/POACEAE 933 12. Pedicels
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Elymus repens [HI1] Elymus smithii
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Eragrostis japonica [USB] ILLUSTRAT
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ERIONEURON/POACEAE 945 Shaw and Sme
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� � � ILLUSTRATED FLORA OF EA
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FESTUCA/POACEAE 951 rachilla glabro
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GYMNOPOGON/POACEAE 955 Glyceria sep
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HILARIA/POACEAE 957 REFERENCES: Eme
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ILLUSTRATED FLORA OF EAST TEXAS Fes
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KOELERIA/POACEAE 963 Andropogoneae,
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Heteropogon melanocarpus [HI1] Hord
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LEPTOCHLOA/POACEAE 967 Leersia oryz
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ILLUSTRATED FLORA OF EAST TEXAS Koe
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ILLUSTRATED FLORA OF EAST TEXAS Hor
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LOLIUM/POACEAE 973 Charmet et al. 1
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Leptochloa nealleyi [USB] Lolium ri
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MICROSTEGIUM/POACEAE 977 AA variabl
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ILLUSTRATED FLORA OF EAST TEXAS Lep
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Lolium temulentum [USB] Melica muti
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MUHLENBERGIA/POACEAE 983 10. Lemma
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MUHLENBERGIA/POACEAE 985 and Prairi
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ILLUSTRATED FLORA OF EAST TEXAS Mon
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ILLUSTRATED FLORA OF EAST TEXAS Mic
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 993 fact that gener
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PANICUM/POACEAE 995 17. Spikelets 2
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Oryza sativa [HI1] Panicum anceps [
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ILLUSTRATED FLORA OF EAST TEXAS Muh
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PANICUM/POACEAE 1001 Panicum hallii
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Panicum trichoides [HI2] ILLUSTRATE
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PASPALUM/POACEAE 1009 5. Spikelets
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PASPALUM/POACEAE 1011 27. Axils of
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Panicum virgatum [RCA] Paspalum acu
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PASPALUM/POACEAE 1019 loam soils; w
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PENNISETUM/POACEAE 1025 unit as in
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Paspalum repens [USB] ILLUSTRATED F
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PHALARIS/POACEAE 1029 landscapes (B
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Paspalum pubiflorum (both vars.) IL
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Pennisetum purpureum [HUT] Penniset
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POA/POACEAE 1035 around awn base; l
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POA/POACEAE 1037 long. Forests, flo
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SACCHARUM/POACEAE 1045 1. Awn of le
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SCHEDONNARDUS/POACEAE 1047 lower gl
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SETARIA/POACEAE 1053 Paspalidium (W
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SETARIA/POACEAE 1055 14. Leaf blade
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Schizachyrium tenerum [HI1, USB] Sc
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SORGHUM/POACEAE 1063 2000) indicate
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SPARTINA/POACEAE 1065 Americas it [
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Sorghastrum elliottii [HI1] Sorghas
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SPOROBOLUS/POACEAE 1071 7. Lemma ap
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STEINCHISMA/POACEAE 1075 Sporobolus
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Sphenopholis obtusata (both vars.)
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Sporobolus neglectus [HI1] Sporobol
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TRICHONEURA/POACEAE 1081 guish it f
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Stenotaphrum secundatum [USB] Trich
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TRISETUM/POACEAE 1089 low areas; ne
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UROCHLOA/POACEAE 1091 AA C4 genus o
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Tridens eragrostoides Tridens flavu
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VULPIA/POACEAE 1097 2. Lemmas (exce
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Page 793 and 794:
Page 795 and 796:
ZIZANIOPSIS/POACEAE 1103 (1999) (po
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Page 799 and 800:
EICHHORNIA/PONTEDERIACEAE 1107 REFE
Page 801 and 802:
Zoysia japonica [HOL] Zoysia pacifi
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POTAMOGETON/POTAMOGETONACEAE 1111 1
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POTAMOGETON/POTAMOGETONACEAE 1113 P
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SMILAX/SMILACACEAE 1119 times woody
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SMILAX/SMILACACEAE 1121 30(-60) mm
Page 815 and 816:
Stuckenia pectinata [MAS] ILLUSTRAT
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THEMIDACEAE 1127 cover for various
Page 821 and 822:
Smilax rotundifolia [JEM] ILLUSTRAT
Page 823 and 824:
TRILLIUM/TRILLIACEAE 1131 Schilling
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TRILLIUM/TRILLIACEAE 1133 taxonomic
Page 827 and 828:
Triantha racemosa [GLE] Trillium lu
Page 829 and 830:
TYPHA/TYPHACEAE 1137 REFERENCES: Wi
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XYRIS/XYRIDACEAE 1139 cone-like spi
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XYRIS/XYRIDACEAE 1141 11. Plant bas
Page 835 and 836:
Sparganium americanum Typha latifol
Page 837 and 838:
Typha domingensis[ BT3] Typha latif
Page 839 and 840:
Xyris drummondii [SID] Xyris fimbri
Page 841 and 842:
Xyris smalliana [SID] ILLUSTRATED F
Page 843:
Xyris caroliniana Xyris difformis (
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