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keys to the vascular plants of east texas - Botanical Research ...

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1138 XYRIDACEAE/XYRIS<br />

as or slightly over<strong>to</strong>pped by <strong>the</strong> leaves. Often very abundant in nutrient-rich ditches, marshes,<br />

s<strong>to</strong>ck tanks, and lake margins, in shallow water or wet ground; nearly throughout TX; s U.S. n <strong>to</strong><br />

VA, NE, WY, and OR. Apr–Jul. During pioneer days, <strong>the</strong> creeping rhizomes “… were eaten, <strong>the</strong><br />

abundant pollen was mixed with flour for <strong>the</strong> making <strong>of</strong> pancakes, and <strong>the</strong> young female inflorescences<br />

were boiled and eaten like miniature roasting ears” (Crosswhite 1980).<br />

Typha latifolia L., (broad-leaved), COMMON CAT-TAIL, BROAD-LEAF CAT-TAIL, TULE ESPADILLA. Plant<br />

<strong>to</strong> ca. 3 m tall; leaf blades 1–2.3(–2.9) cm wide (fresh), 0.5–1.6(–2) cm wide (dry). Ditches,<br />

marshes, s<strong>to</strong>ck tanks, and lake margins, in shallow water or wet ground; throughout much <strong>of</strong><br />

TX; throughout most <strong>of</strong> Canada and <strong>the</strong> U.S. Apr–Jun. These rhizoma<strong>to</strong>us <strong>plants</strong> <strong>of</strong>ten spread <strong>to</strong><br />

form large stands. m/306<br />

Ty pha angustifolia L., (narrow-leaved), NARROW-LEAF CAT-TAIL, is similar <strong>to</strong> T. domingensis but<br />

can be distinguished by <strong>the</strong> inflorescences much over<strong>to</strong>pped by <strong>the</strong> leaves and by <strong>the</strong> characters<br />

in <strong>the</strong> following key. This apparently introduced species (from Europe—Stuckey &<br />

Salamon 1987) was cited for s TX by Correll and Johns<strong>to</strong>n (1970) and for <strong>the</strong> Pineywoods by<br />

Hatch et al. (1990). Jones et al. (1997) synonymized T. angustifolia with T. domingensis, while<br />

Kartesz (1999) indicated that T. angustifolia does not occur in TX. The recent treatment <strong>of</strong><br />

Typha for Flora <strong>of</strong> North America (Smith 2000) indicated that T. angustifolia is a distinct species,<br />

but it is mainly nor<strong>the</strong>rn, does not occur in TX, and only reaches OK sporadically. Smith<br />

(2000) separated T. angustifolia and T. domingensis as follows: I<br />

1. Mucilage glands present on adaxial surface <strong>of</strong> all <strong>of</strong> sheath and usually about 1–10 cm <strong>of</strong> adjacent<br />

blade; pistillate bracteole tips much paler than <strong>to</strong> about same color as stigmas, straw-colored <strong>to</strong><br />

light brown, mostly acute <strong>to</strong> acuminate, in mature spikes exceeding pistil hairs; pistil hair tips<br />

colorless <strong>to</strong> usually orangish (or slightly brownish in hybrids), not evidently enlarged, or <strong>of</strong>ten<br />

with 1 subapical, orange, swollen cell evident at 20–30 �; pistillate spikes bright cinnamon- <strong>to</strong><br />

orange-brown; leaf sheath summits tapered <strong>to</strong> blade or sometimes with membranous auricles<br />

_____________________________________________________________________________ T. domingensis<br />

1. Mucilage glands absent from adaxial surface <strong>of</strong> blade and generally from central part <strong>of</strong> sheath<br />

near sheath summit; pistillate bracteole tips darker than (or as dark as) stigmas, very dark <strong>to</strong><br />

medium brown, rounded (<strong>to</strong> acute), in mature spikes about equaling pistil hairs; pistil hair tips<br />

medium brown, distinctly enlarged at 10–20 � magnification; pistillate spikes medium <strong>to</strong> dark<br />

brown; leaf sheath summits with membranous auricles (<strong>the</strong>se <strong>of</strong>ten disintegrating late in season)<br />

______________________________________________________________________________ T. angustifolia<br />

XYRIDACEAE C. Agardh<br />

YELLOW-EYED-GRASS FAMILY<br />

AA small (over 300 species in 5 genera—Kral 2000a; Kral, pers. comm.) family <strong>of</strong> mainly tropical<br />

and warm area herbs with a few in temperate regions; <strong>the</strong>y usually occur in wet acidic habitats.<br />

The family seems <strong>to</strong> have a close relationship with Eriocaulaceae (Linder & Kellogg 1995;<br />

Stützel 1998; Givnish et al. 1999; Reveal & Pires 2002) or possibly Mayacaceae (Davis et al.<br />

2004). (subclass Commelinidae—Cronquist; order Poales—APG II)<br />

FAMILY RECOGNITION IN THE FIELD: grass-like or rush-like, moist or wet area herbs with basal<br />

leaves and long naked flowering stalks terminated by small, head-like or cone-like spikes with<br />

conspicuous brownish bracts subtending <strong>the</strong> usually yellow flowers.<br />

REFERENCES: Kral 1979a, 1983, 1992, 2000a; Dahlgren et al. 1985; Rudall & Sajo 1999.<br />

XYRIS L. YELLOW-EYED-GRASS<br />

Perennial (except X. jupicai), tufted or solitary, grass- or rush-like scapose herbs; leaves basal,<br />

linear <strong>to</strong> filiform, 2-ranked, equitant; flowering stalks (= scapes) terminated by a head-like or

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