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keys to the vascular plants of east texas - Botanical Research ...

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1108 PONTEDERIACEAE/HETERANTHERA<br />

HETERANTHERA Ruíz & Pav. MUD-PLANTAIN<br />

Annual or perennial herbs submersed or rooted in mud; leaves linear, ribbon- or grass-like,<br />

without a distinct blade, or with ovate <strong>to</strong> elliptic or elliptic-lanceolate blade; flowers solitary (in<br />

East TX species), from a spa<strong>the</strong>; perianth � radially symmetrical, salverform; fruit a manyseeded<br />

capsule.<br />

AA genus <strong>of</strong> 12 species (Horn 2002) <strong>of</strong> tropical and warm areas <strong>of</strong> Africa and <strong>the</strong> Americas n<br />

<strong>to</strong> North America. (Greek: hetera, different, and an<strong>the</strong>ra, an<strong>the</strong>r, from <strong>the</strong> dissimilar an<strong>the</strong>rs <strong>of</strong><br />

<strong>the</strong> first described species)<br />

REFERENCES: Horn 1985, 1988.<br />

1. Leaves all sessile, linear, grass-like, pellucid (= � clear, almost transparent); flowers pale yellow;<br />

<strong>plants</strong> usually completely submersed except for flowers (can become emersed if water levels<br />

drop); spa<strong>the</strong>s sessile in axils <strong>of</strong> leaves ___________________________________________________ H. dubia<br />

1. Leaves (at l<strong>east</strong> some) petiolate, with an expanded, oblong <strong>to</strong> ovate, thickish blade; flowers light<br />

blue <strong>to</strong> purplish blue <strong>to</strong> white; <strong>plants</strong> rooted in mud, <strong>the</strong> leaves forming emersed rosettes or <strong>the</strong><br />

leaves floating at <strong>the</strong> water surface; spa<strong>the</strong>s peduncled ____________________________________ H. limosa<br />

Heteran<strong>the</strong>ra dubia (Jacq.) MacMill., (dubious), GRASS-LEAF MUD-PLANTAIN, WATER STAR-GRASS.<br />

Perennial; petiolate leaves absent; flowers usually exposed at or above <strong>the</strong> water surface; stamens<br />

all alike; an<strong>the</strong>rs coiled with age. Small streams, quiet waters, and ditches, growing most<br />

extensively in alkaline waters (Moyle 1945; Horn 2002); Comal, Travis (BRIT), Brazos (TAMU),<br />

Bastrop (TEX), Dallas, and Liberty (Turner et al. 2003) cos.; scattered in TX, but mainly c and s<br />

parts <strong>of</strong> <strong>the</strong> state; nearly throughout <strong>the</strong> U.S. and se Canada. Apr–Jun. Flowers are reported <strong>to</strong> open<br />

within 2 hours after dawn and <strong>to</strong> wilt by dusk (Horn 2002). [H. liebmannii (Buchneau) Shinners]<br />

Heteran<strong>the</strong>ra limosa (Sw.) Willd., (<strong>of</strong> muddy places), BLUE MUD-PLANTAIN. Annual 6–25 cm tall,<br />

tufted, becoming rhizoma<strong>to</strong>us; both sessile and petiolate leaves present; sessile leaves forming a<br />

basal rosette; petiolate leaves with petioles 2–12 cm long; flowering stem with a terminal<br />

sheathing spa<strong>the</strong>; perianth light blue <strong>to</strong> purplish blue <strong>to</strong> white, with 3 <strong>of</strong> <strong>the</strong> lobes yellow at<br />

base; stamens dimorphic, 2 with short yellow an<strong>the</strong>rs, <strong>the</strong> third with an elongate, light blue or<br />

yellow an<strong>the</strong>r; an<strong>the</strong>rs not coiled. Shallow water or wet places, including pond margins and<br />

ditches; scattered throughout much <strong>of</strong> TX but <strong>of</strong>ten inconspicuous and rarely collected; widespread<br />

in <strong>the</strong> c U.S. from KY and TN w <strong>to</strong> SD and AZ, also CA and FL. Jun–Oct. Flowers are reported<br />

<strong>to</strong> open within 1 hour after dawn and <strong>to</strong> wilt by midday (Horn 2002).<br />

Heteran<strong>the</strong>ra rotundifolia (Kunth) Griseb., (round-leaved), ROUND-LEAF MUD-PLANTAIN, is<br />

known from w TX and, according <strong>to</strong> C. Horn (pers. comm.), could possibly be found in <strong>the</strong> very<br />

w part <strong>of</strong> East TX. Horn (2002) distinguished this species from <strong>the</strong> similar H. limosa as follows:<br />

1. Vegetative stems commonly elongating unless plant emersed from early age; petiolate leaf blade<br />

round <strong>to</strong> oblong, base cordate <strong>to</strong> truncate; distal central perianth limb lobe with lateral flanges<br />

_____________________________________________________________________________ H. rotundifolia<br />

1. Vegetative stems elongating only on <strong>plants</strong> in over 5 cm <strong>of</strong> water; petiolate leaf blade oblong <strong>to</strong><br />

ovate, base truncate <strong>to</strong> cuneate; distal central perianth limb lobe without lateral flanges __________ H. limosa<br />

PONTEDERIA L. PICKEREL-WEED<br />

AA New World genus <strong>of</strong> 6 species (Cook 1998b; Horn 2002) occurring from Canada <strong>to</strong> Argentina.<br />

(Named for Guilio Pontedera, 1688–1757, Italian botanist and pr<strong>of</strong>essor at Padua)<br />

REFERENCES: Fernald 1925; Ornduff 1966; Lowden 1973; Price & Barrett 1982; Horn 1987; Horn<br />

& Haynes 1987.<br />

Pontederia cordata L., (cordate, heart-shaped), PICKEREL-WEED, WAMPEE. Emergent perennial<br />

rooted in mud, 40–80(–120) cm tall, from thick, short-rhizoma<strong>to</strong>us base; leaves both sessile and

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