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1106 PONTEDERIACEAE<br />

(2003) also mapped several counties in <strong>the</strong> Gulf Prairies and Marshes; no county distribution<br />

map is provided; se Canada (Ont.) and widely scattered in <strong>the</strong> U.S. Native <strong>of</strong> Japan. [Z. matrella var.<br />

japonica (Steud.) Sasaki] This species is used for lawns and playing fields (Watson & Dallwitz<br />

1992). According <strong>to</strong> Anderson (2003), this was “<strong>the</strong> first species <strong>of</strong> Zoysia introduced in<strong>to</strong> cultivation<br />

in <strong>the</strong> United States, with <strong>the</strong> introduction <strong>of</strong> <strong>the</strong> cultivar ‘Meyer’ in <strong>the</strong> 1950s. It is <strong>the</strong> most<br />

cold-<strong>to</strong>lerant and coarsely textured <strong>of</strong> <strong>the</strong> three species … and is <strong>the</strong> only species that is currently<br />

available as seed in <strong>the</strong> United States. The o<strong>the</strong>r two species treated here can be established from<br />

seed, but <strong>the</strong>re are currently no commercial sources <strong>of</strong> ei<strong>the</strong>r one in <strong>the</strong> United States.” I<br />

Zoysia matrella (L.) Merr., (Derivation unknown, but possibly from an Urdu word mehtar, headman,<br />

because <strong>the</strong> original author gives <strong>the</strong> location <strong>of</strong> <strong>the</strong> type as from Malabar, India), MANILA<br />

TEMPLE GRASS, MANILA GRASS. Leaf blades <strong>to</strong> 7 cm long, weakly <strong>to</strong> strongly involute; inflorescence<br />

1–4 cm long; spikelets 2.1–3.2 mm long, unawned or with awn <strong>to</strong> 1 mm long. Persisting or<br />

possibly escaping from lawns; included based on indication by S. Anderson (pers. comm.) that<br />

it possibly occurs in <strong>the</strong> area; no specimens have been seen and we are not aware <strong>of</strong> a county<br />

record; no county distribution map is provided; FL, NM, and possibly TX. Native <strong>of</strong> <strong>the</strong> coasts <strong>of</strong><br />

<strong>the</strong> Indian Ocean. [Agrostis matrella L., Z. pungens Willd.] This species is considered by some authorities<br />

<strong>to</strong> be a significant weed species (Watson & Dallwitz 1992). According <strong>to</strong> Anderson<br />

(2003), “Many <strong>of</strong> <strong>the</strong> Zoysia lawn grasses grown in <strong>the</strong> sou<strong>the</strong>rn and <strong>east</strong>ern United States are<br />

derived from hybrids between Z. matrella and Z. pacifica or Z. japonica, and have retained<br />

many <strong>of</strong> <strong>the</strong> characteristics <strong>of</strong> Z. matrella.” I<br />

Zoysia pacifica (Goudswaard) M. Hotta & Kuroki, (<strong>of</strong> <strong>the</strong> Pacific), MASCARENE GRASS,<br />

MASCARENE TEMPLE GRASS, KOREAN VELVET GRASS. Leaf blades 3 cm or less long, involute <strong>to</strong><br />

strongly convolute (<strong>the</strong> margins overlapping); inflorescence 0.4–2 cm long; spikelets 2.2–2.9<br />

mm long, 0.5–0.8 mm wide, lanceolate <strong>to</strong> linear, unawned or with awn <strong>to</strong> 0.5 mm. Persisting or<br />

possibly escaping from lawns; included for East TX based on indication by S. Anderson (pers.<br />

comm.) that it possibly occurs in <strong>the</strong> area; no specimens have been seen and we are not aware <strong>of</strong><br />

a county record; no county distribution map is provided; CA, FL, and possibly TX. Native <strong>of</strong><br />

coasts <strong>of</strong> <strong>the</strong> w Pacific Ocean. [Z. matrella var. pacifica Goudswaard, Z. tenuifolia <strong>of</strong> authors not<br />

Willd. ex Thiele] This is <strong>the</strong> l<strong>east</strong> cold-<strong>to</strong>lerant <strong>of</strong> <strong>the</strong> three species included here and is not a<br />

common lawn grass in <strong>the</strong> U.S. (Anderson 2003). “The cultivar ‘Cashmere’ has many <strong>of</strong> <strong>the</strong><br />

characteristics <strong>of</strong> Z. pacifica; it is probably derived from a hybrid between Z. matrella and Z.<br />

pacifica (Anderson 2003).” I<br />

PONTEDERIACEAE Kunth<br />

WATER-HYACINTH OR PICKEREL-WEED FAMILY<br />

Glabrous perennial or annual herbs; leaves basal or alternate, simple, entire, both sessile and<br />

petiolate leaves <strong>of</strong>ten present; flowers solitary or in spikes, <strong>the</strong> inflorescence subtended by a<br />

spa<strong>the</strong>-like bract; perianth parts united proximally <strong>to</strong> form a very slender basal tube; perianth<br />

lobes 6, one differing slightly or greatly in size, shape, or coloration; stamens 3 or 6; pistil 1;<br />

ovary superior; fruit a many-seeded capsule or a 1-seeded utricle.<br />

AA small (ca. 30 species in 6 genera—Horn 2002) family <strong>of</strong> freshwater aquatic herbs <strong>of</strong> tropical<br />

and warm areas nearly worldwide, but especially in <strong>the</strong> Americas, with a few in <strong>the</strong> n temperate<br />

zone. Some are problematic weeds; a number are cultivated as ornamentals. The family<br />

appears related <strong>to</strong> such familes as <strong>the</strong> Hamemodoraceae and Commelinaceae (Chase et al.<br />

2000; Reveal & Pires 2002). (subclass Liliidae—Cronquist; order Commelinales—APG II)<br />

FAMILY RECOGNITION IN THE FIELD: rooted or free-floating, wet area or aquatic herbs; flowers<br />

<strong>of</strong>ten conspicuous, <strong>the</strong> perianth petaloid; inflorescence a solitary flower or a spike, subtended<br />

by a spa<strong>the</strong>-like bract.

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