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944 POACEAE/EREMOCHLOA<br />

widespread in TX; c U.S. from MI s <strong>to</strong> MS w <strong>to</strong> WY and NM, also NY and VT. Jul–Dec., sporadically<br />

in <strong>the</strong> spring. [E. pilifera Scheele, E. trichodes var. pilifera (Scheele) Fernald] Hatch (2002)<br />

recognized var. pilifera. However, we have not been able <strong>to</strong> find consistent differences and are<br />

thus following Correll and Johns<strong>to</strong>n (1970), Gould (1975b), and Peterson (2003a) in not recognizing<br />

varieties. According <strong>to</strong> Peterson (2003), this species is sometimes used as an ornamental.<br />

EREMOCHLOA Büse CENTIPEDE GRASS<br />

AA genus <strong>of</strong> 11 species (Buitenhuis & Veldkamp 2001; Thieret 2003d) native from<br />

Indomalesia (India <strong>to</strong> New Guinea) <strong>to</strong> Australia. It is considered by some authorities (e.g., Clay<strong>to</strong>n<br />

& Renvoize 1986) <strong>to</strong> be related <strong>to</strong> Coelorachis. The solitary, terminal, 1-sided, spike-like<br />

raceme distinguishes this genus from all o<strong>the</strong>r members <strong>of</strong> <strong>the</strong> tribe Andropogoneae (Bor 1952b).<br />

Like all members <strong>of</strong> <strong>the</strong> Andropogoneae, Eremochloa is characterized by C4 pho<strong>to</strong>syn<strong>the</strong>sis<br />

(Kellogg 2000a). (Greek: eremos, desert, and chloa, grass—Bor 1952b) (subfamily Panicoideae,<br />

tribe Andropogoneae)<br />

REFERENCES: Bor 1952b; Buitenhuis & Veldkamp 2001; Thieret 2003d.<br />

Eremochloa ophiuroides (Munro) Hack., (snake-tailed), CENTIPEDE GRASS. Mat-forming s<strong>to</strong>loniferous<br />

perennial; culms erect, <strong>to</strong> 35 cm tall; leaf sheaths sharply keeled; leaf blades (lower ones)<br />

1–5 mm wide, <strong>the</strong> upper ones reduced or lacking; inflorescence a solitary, terminal, spike-like<br />

raceme ca. 2–6 cm long and ca. 1.5–2.5 mm wide (pencil-like but much smaller in diam.), <strong>the</strong><br />

spikelets closely appressed and overlapping scale-fashion along 1 side; spikelets awnless, glabrous,<br />

in pairs, one sessile, one pedicellate; disarticulation at base <strong>of</strong> sessile spikelet so that associated<br />

pedicel and section <strong>of</strong> inflorescence axis fall with <strong>the</strong> sessile spikelet; pedicelled spikelet<br />

usually much reduced (represented by a bristle) or absent (only � flattened pedicel remaining);<br />

sessile spikelets dorsally compressed, 3–4 mm long, 2-flowered, <strong>the</strong> lower floret staminate, <strong>the</strong><br />

upper floret fertile; lower glume winged near apex, making <strong>the</strong> glume � truncate apically, <strong>the</strong><br />

keel with 1–several hook-like spines 0.2–0.3 mm long near base. Roadsides, parking lots, lawns;<br />

Cherokee (BRIT), Angelina, Liberty, New<strong>to</strong>n (SBSC), Leon (Gould 1975b), and Brazos (Turner et<br />

al. 2003) cos.; according <strong>to</strong> Gould (1975b) this species is an occasional escape from experimental<br />

plantings, but we are unaware <strong>of</strong> o<strong>the</strong>r TX localities; se U.S. from NC s <strong>to</strong> FL w <strong>to</strong> AR and TX,<br />

also MA. Mainly spring–fall. Native <strong>of</strong> e Asia. [Ischaemum ophiuroides Munro] This species is<br />

used for erosion control and as a lawn grass (Mabberley 1997), particularly in <strong>the</strong> tropics (Clay<strong>to</strong>n<br />

& Renvoize 1986). Its use as a lawn grass in <strong>the</strong> sou<strong>the</strong>astern United States dates from ca.<br />

1920 (Thieret 2003d). The extremely slender, spike-like inflorescences superficially resemble<br />

those <strong>of</strong> Coelorachis (JOINT-TAIL). The common name is in reference <strong>to</strong> <strong>the</strong> appearance <strong>of</strong> <strong>the</strong><br />

leafy s<strong>to</strong>lons (Thieret 2003d). I<br />

ERIOCHLOA Kunth CUP GRASS<br />

Annuals or perennials; ligule <strong>of</strong> hairs <strong>to</strong> 1–2 mm long; panicle slender, <strong>of</strong> erect, appressed, spikelike<br />

racemes along a main axis; spikelets each with a small (� 0.5 mm long) cup-like collar or<br />

ring (called <strong>the</strong> callus) just under <strong>the</strong> base, solitary or paired, 2-flowered, <strong>the</strong> lower staminate or<br />

neuter, <strong>the</strong> upper perfect; disarticulation at base <strong>of</strong> spikelets; lower glume absent (see note in<br />

synopsis below); upper glume and lemma <strong>of</strong> sterile floret similar.<br />

AA C4 genus <strong>of</strong> 20–30 species <strong>of</strong> tropical and warm areas <strong>of</strong> <strong>the</strong> world (Shaw & Smeins 1981;<br />

Shaw et al. 2003). More than two-thirds <strong>of</strong> <strong>the</strong> taxa are found in <strong>the</strong> American southwest and in<br />

<strong>the</strong> central Andes (Shaw & Smeins 1981). Some species are important pasture grasses while<br />

o<strong>the</strong>rs are considered significant weeds (Watson & Dallwitz 1992). The small cup-like structure<br />

below <strong>the</strong> spikelets, known as <strong>the</strong> callus, is quite unique and distinguishes <strong>the</strong> genus from<br />

all o<strong>the</strong>r members <strong>of</strong> its tribe (Shaw & Webster 1987). Its ana<strong>to</strong>mical derivation has been studied<br />

in detail (Shaw & Smeins 1979, 1983; Thompson et al. 1990) with conflicting conclusions.

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