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keys to the vascular plants of east texas - Botanical Research ...

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POA/POACEAE 1035<br />

around awn base; lemma base and rachilla <strong>to</strong>ge<strong>the</strong>r forming a sharp-pointed bearded callus;<br />

palea slightly longer than <strong>the</strong> lemma, longitudinally grooved, <strong>the</strong> apex protruding between <strong>the</strong><br />

lemma margins; caryopsis permanently enclosed within <strong>the</strong> lemma. Woodlands in shade, forest<br />

margins, sandy soils; widespread in Pineywoods and Post Oak Savannah w <strong>to</strong> Limes<strong>to</strong>ne,<br />

Robertson (BRIT), and Van Zandt (Turner et al. 2003) cos.; also n Gulf Prairies and Marshes and<br />

e Edwards Plateau; se Canada (Ont.) and e U.S. from MA s <strong>to</strong> FL w <strong>to</strong> MI and TX. Mar–Jun. [Stipa<br />

avenacea L.] This species can be confused with Nassella leucotricha (TEXAS WINTER GRASS), but<br />

can be distinguished by habitat (Nassella in open disturbed sites) by <strong>the</strong> lemmas (Nassella with<br />

lemmas light brown, with overlapping margins, and apically with a smooth white neck ca. 0.6–<br />

1 mm long), and by <strong>the</strong> ligules (Nassella with ligules 1 mm or less long).<br />

POA L. BLUE GRASS<br />

Annuals or perennials, sometimes with rhizomes; ligule a membrane; leaf blades sometimes<br />

with tips cupped; inflorescences open or contracted panicles; spikelets relatively small, usually<br />

with 2–6(–10) florets, awnless, disarticulating above glumes and between florets; glumes<br />

shorter than rest <strong>of</strong> spikelet; lemmas acute or obtuse, usually keeled on back, <strong>of</strong>ten with long,<br />

kinky hairs at base.<br />

AA C3 genus <strong>of</strong> ca. 575 species, Poa is <strong>the</strong> largest genus <strong>of</strong> grasses (Gillespie & Soreng 2005). It<br />

is widespread but principally <strong>of</strong> temperate and cold areas <strong>of</strong> <strong>the</strong> world, including tropical<br />

mountains. Some are important as fodder grasses and for use in lawns and playing fields; many<br />

are important as pasture species, while a number are significant weeds (Watson & Dallwitz<br />

1992). The genus approaches Festuca (lemmas rounded on back), but differs in its keeled lemma<br />

(Clay<strong>to</strong>n & Renvoize 1986). Poa is morphologically ra<strong>the</strong>r uniform (Clay<strong>to</strong>n & Renvoize 1986),<br />

and <strong>the</strong>re is apparently little agreement on subdivisions within <strong>the</strong> genus (Tucker 1996). However,<br />

recent molecular research (e.g., Gillespie & Soreng 2005) has begun <strong>to</strong> clarify <strong>the</strong> broad<br />

phylogenetic structure within <strong>the</strong> genus. It is a taxonomically difficult group because <strong>of</strong> <strong>the</strong><br />

widespread occurrence <strong>of</strong> polyploidy, apomixis, hybridization, and introgression (Clay<strong>to</strong>n &<br />

Renvoize 1986; Soreng & Kellogg ined.). Some <strong>of</strong> <strong>the</strong> species have variable numbers <strong>of</strong> chromosomes,<br />

including aneuploid series (Tucker 1996). (Greek: poa, ancient name for grass or fodder)<br />

(subfamily Pooideae, tribe Poeae)<br />

REFERENCES: Marsh 1952; Hutchinson & Seymour 1982; Soreng 1990, 1998; Tucker 1996; Soreng<br />

& Barrie 1999; Soreng & Kellogg ined; Gillespie & Soreng 2005.<br />

1. Florets developing in<strong>to</strong> bulbils (<strong>of</strong>ten dark purple at base), <strong>the</strong> lemmas prolonged <strong>to</strong> 5–15 mm<br />

long as if sprouting; culms swollen at base; rare in TX, known only from Den<strong>to</strong>n Co. near w margin<br />

<strong>of</strong> East TX _____________________________________________________________________ P. bulbosa<br />

1. Florets not developing in<strong>to</strong> bulbils; culms not swollen at base; including species widespread<br />

and abundant in East TX.<br />

2. Plants annual, 3–45 cm tall, without rhizomes; keel <strong>of</strong> glumes smooth (P. annua) or minutely<br />

scabrous-ciliate (P. bigelovii and P. chapmaniana).<br />

3. Branches <strong>of</strong> inflorescence appressed-erect, <strong>the</strong> inflorescence thus narrow and contracted,<br />

usually � 1 cm wide; species known in East TX only from Travis Co. on extreme w margin <strong>of</strong><br />

area _______________________________________________________________________ P. bigelovii<br />

3. Branches <strong>of</strong> inflorescence spreading (at l<strong>east</strong> <strong>the</strong> lower ones), <strong>the</strong> inflorescence thus not<br />

narrow, usually � 1 cm wide; including species widespread in East TX.<br />

4. Lemmas with long, kinky, cobwebby hairs at base, with 3 strong and 2 faint veins; an<strong>the</strong>rs<br />

0.1–0.3 mm long; species rare in East TX ________________________________ P. chapmaniana<br />

4. Lemmas usually without long hairs at base (but long-pubescent on margins and keel),<br />

with 5 strong veins; an<strong>the</strong>rs 0.5–1.3 mm long; species widespread and abundant in East<br />

TX ________________________________________________________________________ P. annua

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