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keys to the vascular plants of east texas - Botanical Research ...

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ANDROPOGON/POACEAE<br />

821<br />

colonizing early successional or permanently disturbed sites (Campbell 1982). (Greek: aner<br />

(andr), man, and pogon, beard, referring <strong>to</strong> <strong>the</strong> hairy pedicels <strong>of</strong> <strong>the</strong> staminate spikelets) (subfamily<br />

Panicoideae, tribe Andropogoneae)<br />

REFERENCES: Gould 1957a, 1967; Campbell 1982, 1983a, 1986; Barnes 1986; Wipff 1996b;<br />

Gustafson et al. 1999; Campbell 2003.<br />

1. Sessile spikelets 7–11 mm long; terminal inflorescence conspicuously exserted beyond bracts,<br />

not woolly in appearance; pedicelled spikelets large, well-developed, similar <strong>to</strong> sessile spikelets<br />

except awnless ___________________________________________________________________ A. gerardii<br />

1. Sessile spikelets 7 mm or less long; inflorescences ei<strong>the</strong>r not conspicuously exserted beyond<br />

bracts OR if exserted <strong>the</strong>n appearing woolly; pedicelled spikelets rudimentary, vestigial, or represented<br />

by <strong>the</strong> pedicel only.<br />

2. Inflorescence with 2 relatively stiff and usually straight branches, <strong>the</strong> spikelets and hairs usually<br />

so dense that <strong>the</strong> branch axes are not easily visible; sessile spikelets 5–7 mm long; inflorescences<br />

<strong>of</strong>ten conspicuously exserted beyond bracts _______________________________ A. ternarius<br />

2. Inflorescence with 2–5 slender, delicate and flexuous branches, <strong>the</strong> spikelets and hairs <strong>of</strong>ten not<br />

as dense, <strong>the</strong> branch axes thus <strong>of</strong>ten visible, sometimes easily so; sessile spikelets 5 mm long or<br />

less; inflorescences usually not conspicuously exserted OR sometimes exserted in A. gyrans.<br />

3. Upper sheathing bract <strong>of</strong> inflorescence conspicuously inflated and spa<strong>the</strong>-like; lemma awns<br />

with a loose spiral twist near base; leaf blades 0.8–5 mm wide (typically ca. 2.5 mm); culms<br />

with tufts <strong>of</strong> long hairs just below sheathing bract <strong>of</strong> inflorescence; species rare in East TX ______ A. gyrans<br />

3. Upper sheathing bract <strong>of</strong> inflorescence not inflated; lemma awns straight or nearly so; leaf<br />

blades 1.7–9.5 mm wide (usually � 3 mm); culms glabrous OR with tufts <strong>of</strong> long hairs just<br />

below sheathing bract <strong>of</strong> inflorescence; species extremely widespread and abundant in<br />

East TX.<br />

4. Inflorescences crowded apically, <strong>to</strong>ge<strong>the</strong>r appearing broom-like; culms with tuft <strong>of</strong> long<br />

hairs below sheathing bract <strong>of</strong> inflorescence; leaf sheaths usually scabrous, sometimes<br />

smooth; leaf blades 13–109 cm long; ligule 0.6–2.2 mm long _____________________ A. glomeratus<br />

4. Inflorescences numerous but not clustered apically, not broom-like; culms glabrous below<br />

sheathing bract <strong>of</strong> inflorescence; leaf sheaths usually smooth, rarely scabrous; leaf<br />

blades 11–52 cm long; ligule 0.2–1 mm long ___________________________________ A. virginicus<br />

Andropogon gerardii Vitman, (for Louis Gérard, 1733–1819, French botanist). Plant <strong>to</strong> 2 m tall,<br />

essentially glabrous, <strong>of</strong>ten glaucous; inflorescences <strong>of</strong> 2–7 digitately arranged branches, <strong>the</strong><br />

branches 4–11 cm long; sessile spikelets usually scabrous, <strong>of</strong>ten glaucous; upper lemma <strong>of</strong><br />

sessile spikelet awned (sometimes awnless in subsp. hallii); pedicellate spikelets about as large<br />

as sessile, awnless. Aug–Nov. This polyploid species has chromosome numbers <strong>of</strong> both 2n = 6x =<br />

60 and 2n = 9x = 90; <strong>the</strong> <strong>plants</strong> are virtually indistinguishable morphologically (Norrmann et<br />

al. 1997). Hybridization between <strong>the</strong>se two main cy<strong>to</strong>types is possible, with <strong>the</strong> resulting individuals<br />

having a range <strong>of</strong> chromosome numbers (Norrmann & Keeler 2003). The reproductive<br />

biology <strong>of</strong> this species was studied by McKone et al. (1998) and was found <strong>to</strong> be very unusual in<br />

allocating more resources <strong>to</strong> male reproduction (pollen production) than <strong>to</strong> female reproduction<br />

(seed production). The following key <strong>to</strong> subspecies is modified from Su<strong>the</strong>rland (1986) and<br />

Wipff (1996b).<br />

1. Upper lemma <strong>of</strong> sessile spikelet with an awn (7.5–)10–25 mm long; rhizomes short or absent,<br />

with internodes usually 2 mm or less in length; an<strong>the</strong>rs usually � 3.8 mm long; ligule 0.4–2.5 mm<br />

long ________________________________________________________________________ subsp. gerardii<br />

1. Upper lemma <strong>of</strong> sessile spikelet awnless or with an awn <strong>to</strong> 8(–11) mm long; rhizomes welldeveloped,<br />

creeping, with internodes <strong>of</strong>ten exceeding 20 mm in length; an<strong>the</strong>rs usually � 3.8<br />

mm long; ligule (0.9–)3–4.5 mm long _________________________________________________ subsp. hallii

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