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keys to the vascular plants of east texas - Botanical Research ...

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1014 POACEAE/PASPALUM<br />

Abundant weed in disturbed sites, lawns, and roadsides; throughout TX; widespread in s 1/2 <strong>of</strong><br />

<strong>the</strong> U.S., also OR. May–Nov. Native <strong>of</strong> South America (Brazil and Argentina). This species is reported<br />

<strong>to</strong> be commercially important as a forage crop in <strong>the</strong> tropics (Yatskievych 1999), but it is<br />

sometimes a troublesome weed (Clay<strong>to</strong>n & Renvoize 1986). Ergot alkaloids can be present—see<br />

generic synopsis. While almost all grasses are wind-pollinated, this grass is unusual in sometimes<br />

being pollinated at l<strong>east</strong> partially by halictid bees (Adams et al. 1981). � I<br />

Paspalum dissectum (L.) L., (dissected, deeply cut), MUDBANK PASPALUM, MUDBANK CROWN GRASS.<br />

Mat- or clump-forming perennial without conspicuous rhizomes; culms 15–60 cm long; leaf<br />

sheaths glabrous; ligule 1.5–2.5 mm long; inflorescence branches 2–6, 1–3(–5.3) cm long, <strong>the</strong><br />

branch axes broadly winged, sometimes almost leaf-like, as wide or wider than <strong>the</strong> rows <strong>of</strong><br />

spikelets, <strong>the</strong> wing margins sometimes even slightly wrapping around <strong>the</strong> spikelets; spikelets<br />

solitary (with nei<strong>the</strong>r aborted spikelet nor pedicel present immediately beside or below fertile<br />

spikelet), in 2 rows, 1.7–2.1(–2.3) mm long; lower glume absent; upper glume minutely pubescent;<br />

lemma <strong>of</strong> sterile floret glabrous. Lake shores, ditches, sandy soils; Leon (BRIT), Harris<br />

(SBSC), Guadalupe, Montgomery (Morrone et al. 1996), Angelina, Bastrop, Caldwell, Gonzales,<br />

and Nacogdoches (Turner et al. 2003) cos.; also Gulf Prairies and Marshes; e U.S. from NJ s <strong>to</strong> FL<br />

w <strong>to</strong> KS and TX. Jul–Nov. [Panicum dissectum L.] This species can have cleis<strong>to</strong>gamous spikelets<br />

with very small an<strong>the</strong>rs (ca. 0.3 mm long); such spikelets do not open <strong>to</strong> expose <strong>the</strong> an<strong>the</strong>rs and<br />

stigmas and are thus self-pollinated (Morrone et al. 1996).<br />

Paspalum distichum L., (two-spiked), KNOT GRASS, ETERNITY GRASS, JOINT GRASS, FORT THOMPSON<br />

GRASS, JOINTED CROWN GRASS, THOMPSON GRASS. Perennial 10–65 cm tall, with trailing and rooting<br />

culms (s<strong>to</strong>lons) sometimes several meters long; leaf sheaths usually glabrous or pilose;<br />

ligule 0.6–2 mm long; inflorescence branches 2, occasionally 3–4, <strong>the</strong>se usually paired or one<br />

slightly below o<strong>the</strong>r, 1–5(–7) cm long, <strong>the</strong> branch axes only slightly winged; spikelets 2.4–3(–3.2)<br />

mm long, mostly solitary, usually in 2 rows; lower glume <strong>of</strong>ten present, reduced; upper glume<br />

usually minutely pubescent; lemma <strong>of</strong> sterile floret glabrous. Moist or wet areas along ponds,<br />

lakes, streams, and ditches; throughout TX; s 1/2 <strong>of</strong> <strong>the</strong> U.S., also NJ, OR, and WA. Jun–Oct.<br />

[Digitaria paspaloides Michx., P. distichum var. indutum Shinners, P. paspaloides (Michx.)<br />

Scribn.] While some authorities (e.g., Jones et al. 1997) recognize var. indutum (known only<br />

from Dallas Co.—Gould 1975b) with leaf sheaths pubescent, we are following Allred (1982),<br />

Allen (1992b), Kartesz (1999), and Allen and Hall (2003) in not recognizing infraspecific taxa.<br />

Larry Brown (pers. comm.) notes that <strong>the</strong> upper glume can be pubescent on some spikelets but<br />

glabrous on o<strong>the</strong>rs (on <strong>the</strong> same plant) and that likewise <strong>the</strong> first glume can be ei<strong>the</strong>r present or<br />

absent—<strong>the</strong>se conditions suggest that <strong>the</strong> species should perhaps be merged with P. vaginatum.<br />

Paspalum floridanum Michx., (<strong>of</strong> Florida), FLORIDA PASPALUM, BIG FLORIDA PASPALUM, BIG<br />

PASPALUM, FLORIDA CROWN GRASS. Rhizoma<strong>to</strong>us perennial; culms erect, usually 1–2.1 m tall; leaf<br />

sheaths and blades nearly or completely glabrous or � densely hirsute; ligule 1–3.3 mm long;<br />

inflorescence branches usually 2–5(–8), 3–13(–17) cm long, <strong>the</strong> branch axes � unwinged; spikelets<br />

usually 3.6–4.8 mm long, mostly paired, in 4 rows, glabrous; lower glume absent. Grasslands<br />

and open woodlands; widespread in e 1/3 <strong>of</strong> TX; e U.S. from NJ s <strong>to</strong> FL w <strong>to</strong> KS and TX.<br />

Aug–Nov. [Paspalum floridanum var. glabratum Engelm. ex Vasey] Some authorities (e.g., Jones<br />

et al. 1997; Yatskievych 1999; Hatch 2002) recognize var. glabratum, based on � glabrous leaf<br />

sheaths and blades (versus with pubescence). However, we are following Allen and Hall (2003)<br />

in not recognizing infraspecific taxa.<br />

Paspalum hartwegianum E. Fourn., (for Theodore Hartweg, 1812–1871, Royal Horticultural Society<br />

collec<strong>to</strong>r in CA and Mexico), HARTWEG’S PASPALUM, HARTWEG’S CROWN GRASS. Tufted perennial<br />

similar <strong>to</strong> P. pubiflorum; inflorescence branches (3–)4–8, 3–7 cm long, <strong>the</strong> branch axes<br />

winged; spikelets (2.2–)2.5–2.9(–3.2) mm long, typically paired, usually in 4 rows on <strong>the</strong> branch<br />

axis; lower glume absent; upper glume and lemma <strong>of</strong> sterile floret pubescent. Ditches, low areas;

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