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keys to the vascular plants of east texas - Botanical Research ...

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ELEOCHARIS/CYPERACEAE<br />

591<br />

43. Scales <strong>of</strong> spikelets usually acute or subacute<br />

apically; tubercles distinctly wider than tall;<br />

culms hollow with complete transverse<br />

septa 2–4 mm apart (<strong>the</strong>se are evident only<br />

upon internal examination <strong>of</strong> culm); species<br />

known in East TX only from Guadalupe Co.<br />

at s margin <strong>of</strong> area _______________________ E. ravenelii<br />

Eleocharis acicularis (L.) Roem. & Schult., (needle-shaped), NEEDLE SPIKE-RUSH, SLENDER SPIKE-<br />

RUSH, LEAST SPIKE-RUSH. Small perennial <strong>of</strong>ten forming mats, with hair-like rhizomes; culms 2–<br />

23(–60) cm tall, rounded <strong>to</strong> slightly flattened, usually with 4–12 angles or ridges; distal leaf<br />

sheath apex membranous, nearly truncate <strong>to</strong> angled on one side <strong>to</strong> a broad point; spikelets usually<br />

2–6 mm long, ovoid <strong>to</strong> lanceoloid or subcylindric, usually with 5–15 scales (and thus flowers);<br />

achenes bluntly trigonous, 0.7–1.1 mm long, with a number <strong>of</strong> distinct ribs or ridges running<br />

lengthwise and many fine horizontal lines between <strong>the</strong> ribs, usually pearly white;<br />

tubercle small, 0.2 mm or less long. Damp soils, <strong>of</strong>ten on receding shorelines, sometimes submerged;<br />

Henderson, Leon (BRIT), Bastrop, Bexar, Fannin, Gonzales, San Jacin<strong>to</strong>, Travis, and<br />

Walker (Turner et al. 2003) cos.; scattered in TX; throughout Canada and <strong>the</strong> U.S. May–Aug,<br />

sporadically <strong>to</strong> Oct. [E. acicularis var. gracilescens Svenson, E. acicularis var. porcata S.G. Sm.; E.<br />

acicularis var. submersa (Nilsson) Svenson] This species is reported <strong>to</strong> be “able <strong>to</strong> grow in acidic<br />

run<strong>of</strong>f from Appalachian coal mines and flourish in streams with pH as low as 2.8” (Tucker 1987).<br />

Yatskievych (1999) reported that mats <strong>of</strong> this species “can grow so densely as <strong>to</strong> inhibit <strong>the</strong> establishment<br />

<strong>of</strong> o<strong>the</strong>r aquatic species. …” “It is abundant and ecologically important throughout much<br />

<strong>of</strong> its range” (Smith et al. 2002). Smith et al. (2002) also noted that “Submerged, usually nonflowering<br />

<strong>plants</strong> are abundant throughout much <strong>of</strong> <strong>the</strong> range <strong>of</strong> <strong>the</strong> species.” The variant, sometimes<br />

recognized as E. acicularis var. porcata, is sometimes mistaken for E. wolfii (Smith 2001, 2002d).<br />

Eleocharis albida Torr., (white), WHITE SPIKE-RUSH. Rhizoma<strong>to</strong>us perennial forming mats; culms<br />

40 cm or less tall; distal leaf sheath apex light-colored marginally, angled <strong>to</strong> a point on one side,<br />

but without an abrupt <strong>to</strong>oth; spikelets 4–12 mm long, oblong <strong>to</strong> oblong-subcylindric, with 20–<br />

100 scales; achenes trigonous, 0.8–1 mm long, ripening dark brown; tubercles 0.2–0.3 mm long.<br />

Marshes, ditches, o<strong>the</strong>r wet areas; Hardin Co. (Turner et al. 2003); mainly Gulf Prairies and<br />

Marshes; se U.S. from MD s <strong>to</strong> FL w <strong>to</strong> TX, also NM. Summer.<br />

Eleocharis atropurpurea (Retz.) J. Presl & C. Presl, (dark purple), PURPLE SPIKE-RUSH. Tufted annual;<br />

culms 2–15(–25) cm tall; distal leaf sheath apex angled on one side <strong>to</strong> a point; spikelets 2–<br />

6(–8) mm long, ellipsoid <strong>to</strong> lance-ovoid, with (10–)40–80(–100) scales; achenes biconvex, 0.3–<br />

0.6 mm long, dark reddish black <strong>to</strong> black at maturity; tubercle tiny, 0.2 mm or less long,<br />

perched on summit <strong>of</strong> achene. Lake and pond margins, ditches, and o<strong>the</strong>r wet areas; Bastrop,<br />

DeWitt, Hays, and Montgomery (Turner et al. 2003) cos.; sparsely scattered elsewhere in TX; sw<br />

Canada and widely scattered in <strong>the</strong> U.S. Jun–Sep. [Scirpus atropurpureus Retz.]<br />

Eleocharis baldwinii (Torr.) Chapm., (for its discoverer, William Baldwin, 1779–1819, Pennsylvania<br />

botanist and physician), BALDWIN’S SPIKE-RUSH. Delicate tufted annual or perennial [?],<br />

s<strong>to</strong>loniferous, mat-forming; culms thread-like, 3–20(–30) cm tall, <strong>of</strong>ten arching; distal leaf<br />

sheath apex angled on one side <strong>to</strong> a broad point; spikelets flattish, 2–7 mm long, with few (2–<br />

4(–9)) scales, <strong>the</strong> spikelets <strong>of</strong>ten proliferating vegetatively; pistillate spikelets <strong>of</strong>ten present at<br />

base <strong>of</strong> plant; scales <strong>of</strong> spikelets � 2-ranked, folded along <strong>the</strong> midrib and thus boat-like;<br />

achenes trigonous, ca. 0.5–0.9 mm long, whitish buffy <strong>to</strong> olive, grayish olive, or brownish olive;<br />

tubercle conic-subulate, 0.2–0.3(–0.4) mm long. Submerged <strong>to</strong> emergent in lakes, ponds, or<br />

o<strong>the</strong>r wet areas; scattered in East TX; se U.S. from NC s <strong>to</strong> FL w <strong>to</strong> AR and TX. End <strong>of</strong> Apr–Oct.<br />

[E. capillacea <strong>of</strong> authors, not Kunth]

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