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PHALARIS/POACEAE<br />

1029<br />

landscapes (Barkworth ined.). � Phalaris arundinacea L., REED CANARY GRASS, a circumpolar<br />

species native <strong>to</strong> <strong>the</strong> n U.S., is weedy and difficult <strong>to</strong> eradicate and is considered a noxious weed<br />

in WA and NC (Kartesz 1999). � Some species <strong>of</strong> Phalaris are potentially <strong>to</strong>xic <strong>to</strong> grazing lives<strong>to</strong>ck,<br />

with <strong>to</strong>xins including alkaloids (Clay 1988; Burrows & Tyrl 2001). (Ancient Greek name<br />

for an undetermined grass) (subfamily Pooideae, tribe Poeae)<br />

REFERENCES: Anderson 1961; Baldini 1995; Tucker 1996; Hatch et al. 2004; Barkworth ined.<br />

1. Inflorescences usually 6–15(–20) cm long; glumes 3–5.5 mm long, with lateral veins scabrous<br />

with 9 or more stiff hairs; <strong>plants</strong> usually 60–170 cm tall; species known from <strong>the</strong> s part <strong>of</strong> East TX<br />

________________________________________________________________________________ P. angusta<br />

1. Inflorescences usually 1.5–7 cm long; glumes 4–10 mm long, with lateral veins glabrous OR scabrous<br />

with 5 or fewer stiff hairs; <strong>plants</strong> usually 25–70(–100) cm tall; including species widespread<br />

in East TX.<br />

2. Glumes usually 6–10 mm long, <strong>the</strong> keel with a conspicuous wing <strong>to</strong> 1 mm wide near <strong>the</strong> tip;<br />

caryopsis (= fruit) 3.9–4.2 mm long; reduced lemmas 2.5–4.5 mm long, more than 1/2 as long<br />

as lemma <strong>of</strong> fertile floret; lemma <strong>of</strong> fertile floret 4.5–6.8 mm long; introduced species rare in<br />

East TX _____________________________________________________________________ P. canariensis<br />

2. Glumes 4.2–6 mm long, <strong>the</strong> keel with only a narrow wing <strong>to</strong> 0.5 mm wide near <strong>the</strong> tip; caryopsis<br />

1.7–2.4 mm long; reduced lemmas 1.5–2.5 mm long, 1/3–1/2 as long as lemma <strong>of</strong> fertile<br />

floret; lemma <strong>of</strong> fertile floret 3–4.7 mm long; native species widespread and common<br />

in East TX ___________________________________________________________________ P. caroliniana<br />

Phalaris angusta Nees ex Trin., (narrow), TIMOTHY CANARY GRASS. Culms usually 60–170 cm tall;<br />

ligule a membrane 3–7 mm long; leaf blades 3–18 cm long, 5–12 mm wide; inflorescences elongate,<br />

6–15(–20) cm long, ca. 8–10 mm wide; glumes ca. 3–5.5 mm long; reduced lemmas 0.5–1.5<br />

mm long; lemma <strong>of</strong> fertile floret (2–)2.3–3.2(–3.8) mm long; caryopsis less than 1.5 mm long.<br />

Ditches and o<strong>the</strong>r open moist areas; Hardin, Harris, Jefferson (BRIT), Brazos, Liberty, and Orange<br />

(Turner et al. 2003) cos. in se part <strong>of</strong> East TX; also Gulf Prairies and Marshes and Blanco<br />

Co. (Turner et al. 2003) in e Edwards Plateau; s 1/4 <strong>of</strong> <strong>the</strong> U.S. from SC s <strong>to</strong> FL w <strong>to</strong> CA, also OR.<br />

Apr–May. This species has an interesting distribution, occurring in both North and South<br />

America; <strong>the</strong>re is some doubt about where it originated (Baldini 1995). The epi<strong>the</strong>t angustata<br />

has sometimes been mistakenly used for this species (e.g., Gould 1975b).<br />

Phalaris canariensis L., (<strong>of</strong> <strong>the</strong> Canary Islands), CANARY GRASS. Culms 25–70(–100) cm tall;<br />

ligule a membrane 2–6 mm long; leaf blades 5–15(–20+) cm long, 3–10 mm wide; inflorescences<br />

short, thick, 1.5–3(–4) cm long, 10–20 mm wide; glumes 6–10 mm long, glabrous or sparsely hispid,<br />

not scabrous, � pale with dark green along <strong>the</strong> lateral veins. Waste areas, probably coming<br />

up from discarded birdseed and not long persisting; Brazos (BRIT), Bell (HPC), Bexar, Hill, Limes<strong>to</strong>ne,<br />

and Nacogdoches (Turner et al. 2003) cos.; widely scattered in TX; throughout most <strong>of</strong><br />

Canada and <strong>the</strong> U.S. Mar–Jun. Native <strong>of</strong> s Europe and <strong>the</strong> Canary Islands. This species is important<br />

as a source <strong>of</strong> commercial bird feed; it is <strong>the</strong> canary seed <strong>of</strong> commerce (Gould 1975b;<br />

Mabberley 1987). I<br />

Phalaris caroliniana Walter, (<strong>of</strong> Carolina), WILD CANARY GRASS, CAROLINA CANARY GRASS,<br />

SOUTHERN CANARY GRASS, MAY GRASS. Culms usually 25–70(–100) cm tall; ligule a membrane 1–<br />

5 mm long; leaf blades 6–15(–20) cm long, 3–10(–13) mm wide; inflorescences cylindrical <strong>to</strong> elliptic<br />

or nearly ovate in outline, usually 7 cm or less long (see note below), 8–13(–20) mm wide;<br />

glumes 4.2–6 mm long, <strong>the</strong> lateral veins glabrous or scabrous with 5 or less spicules. Moist areas,<br />

ravines, disturbed sites, <strong>of</strong>ten in sandy or gravelly soils; throughout TX; s 1/2 <strong>of</strong> <strong>the</strong> U.S.<br />

Apr–Jun. A few aberrant individuals <strong>of</strong> P. caroliniana with elongate inflorescences <strong>to</strong> 12 cm<br />

long (e.g., Whitehouse 15711, Dallas Co., BRIT) have been observed; <strong>the</strong>y are reminiscent <strong>of</strong> P.<br />

angusta, but can be distinguished by <strong>the</strong> o<strong>the</strong>r characters in <strong>the</strong> key <strong>to</strong> species.

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