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470 ARACEAE/PELTANDRA<br />

peduncle <strong>to</strong> rhizome; flowers usually perfect, variable; perianth segments 2–6; stamens 1–6 (1–2<br />

staminodes sometimes present); pistil usually 1; fruit a blue-green, 1-seeded berry ca. 10–20 mm<br />

in diam. Shallow water, moist banks; Polk Co. (BRIT) in <strong>the</strong> Pineywoods; e U.S. from NY s <strong>to</strong> FL<br />

w <strong>to</strong> TX. Apr–Jun. When properly prepared (boiled repeatedly, etc.), <strong>the</strong> rhizome and seeds are<br />

reported <strong>to</strong> be edible (Mabberley 1997), and <strong>the</strong> seeds were eaten by Native Americans (Thompson<br />

2000b). Feb–Jun. Thompson (2000b) suggested that this is probably <strong>the</strong> most distinctive<br />

species <strong>of</strong> Araceae in North America, citing <strong>the</strong> absence <strong>of</strong> a spa<strong>the</strong> and <strong>the</strong> characteristic bluegreen<br />

velvety leaves. The presence/absence <strong>of</strong> a spa<strong>the</strong> in this species is controversial. According<br />

<strong>to</strong> Ray (1988), <strong>the</strong> sheath at <strong>the</strong> base <strong>of</strong> <strong>the</strong> peduncle, with only a bract-like blade, is not<br />

morphologically equivalent <strong>to</strong> <strong>the</strong> spa<strong>the</strong> <strong>of</strong> o<strong>the</strong>r members <strong>of</strong> <strong>the</strong> family; T. Croat (pers. comm.),<br />

however, considers this structure <strong>to</strong> be a spa<strong>the</strong>. This species is sometimes cultivated as an ornamental<br />

(Thompson 2000b). While not <strong>of</strong>ficially designated as such (e.g., TOES 1993; Carr<br />

2002d; Poole et al. 2002), given its limited distribution in <strong>the</strong> state, we consider this species <strong>to</strong> be<br />

<strong>of</strong> conservation concern in TX. � m/293<br />

PELTANDRA Raf. ARROW ARUM<br />

AA genus <strong>of</strong> 2 species endemic <strong>to</strong> e North America (Thompson 2000b); some authorities (e.g.,<br />

Blackwell & Blackwell 1974 [1975]) believe <strong>the</strong>re is only a single variable species. (Greek: pelta,<br />

shield, and andr, man, in reference <strong>to</strong> <strong>the</strong> shield-shaped <strong>to</strong>ps <strong>of</strong> <strong>the</strong> staminate flowers—Thompson<br />

2000b)<br />

REFERENCES: Blake 1912; Goldberg 1941; Wilson 1960; Blackwell & Blackwell 1974 [1975]; Patt et<br />

al. 1995.<br />

Peltandra virginica (L.) Schott, (<strong>of</strong> Virginia), VIRGINIA ARUM, TUCKAHOE, GREEN ARUM, ARROW<br />

ARUM, GREEN ARROW ARUM. Perennial monoecious herb from thick fibrous roots; leaves clustered<br />

on short stems; petioles <strong>to</strong> ca. 0.6(–1) m long; leaf blades highly variable in shape and size,<br />

<strong>to</strong> 50(–59) cm long and ca. 15(–30) cm wide (usually small in comparison <strong>to</strong> those <strong>of</strong><br />

Xanthosoma), with 3 main veins, entire, somewhat glaucous beneath or not glaucous, � sagittate<br />

or hastate, with basal lobes varying from short and rounded <strong>to</strong> long and pointed; scape<br />

nearly equaling <strong>the</strong> petioles in length at flowering time; spa<strong>the</strong> green with pale <strong>to</strong> white or yellow-green<br />

margins, ca. 10–20 cm long, open distally or almost throughout, <strong>the</strong> lower part persistent<br />

and enclosing <strong>the</strong> spadix in fruit; spadix white <strong>to</strong> cream, yellow, or orangish, from<br />

shorter than <strong>to</strong> nearly as long as spa<strong>the</strong>, completely covered by flowers or only <strong>the</strong> apex naked,<br />

<strong>the</strong> lower portion pistillate, <strong>the</strong> upper portion staminate, with sterile flowers between; perianths<br />

absent; staminate flowers with 4–5 connate stamens forming a flat-<strong>to</strong>pped synandrium;<br />

berries ca. 6–18 mm in diam., green <strong>to</strong> brownish or dark purple-green, with 1–2(–4) seeds surrounded<br />

by gelatinous material. Bogs, swamps, and o<strong>the</strong>r wet areas; Cherokee, Henderson,<br />

Hous<strong>to</strong>n, Leon, Robertson, San Augustine, Smith (BRIT), Anderson, Marion, Wood (TAES),<br />

Nacogdoches (ASTC), Bowie (BAYLU), Van Zandt (Kral 1955), Cass, Falls, Madison, and Orange<br />

(Turner et al. 2003) cos., Pineywoods and Post Oak Savannah; also n Gulf Prairies and Marshes;<br />

se Canada and throughout e U.S. w <strong>to</strong> MN and TX, also naturalized in CA and OR. Apr–May. [P.<br />

tharpii F.A. Barkley] Pollination is by a chloropid fly; <strong>the</strong> flies use VIRGINIA ARUM inflorescences<br />

as mating and brood sites, with both adults and developing larvae feeding on pollen (but not<br />

damaging ovules or seeds); this type <strong>of</strong> pollination system has been referred <strong>to</strong> as <strong>the</strong><br />

“androecial brood-site pollination syndrome” (Patt et al. 1995; Thompson 2000b). Fruits and<br />

seeds <strong>of</strong> this species are eaten by wildlife, especially waterfowl. Migra<strong>to</strong>ry birds are thought <strong>to</strong><br />

be an important fac<strong>to</strong>r in dispersal (Thompson 2000b). The roasted starchy roots were formerly<br />

used as food by Native Americans (Mabberley 1997); <strong>the</strong>y are reportedly poisonous until<br />

properly prepared by cooking (Yatskievych 1999). The common name TUCKAHOE is one <strong>of</strong> <strong>the</strong><br />

few Native American plant names still widely used; it was written about by Captain John<br />

Smith (Durant 1976) �

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