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ELEOCHARIS/CYPERACEAE<br />

589<br />

27. Spikelets relatively short, only (1.5–)3–7(–9) mm long even with age; leaf sheaths<br />

and rhizomes usually not purplish red; distal leaf sheaths with apex inflated, loose,<br />

thin-membranous and transparent, <strong>of</strong>ten conspicuously wrinkled, frequently disintegrating;<br />

<strong>plants</strong> 42 cm or less tall _____________________________________ E. flavescens<br />

27. Spikelets relatively long, usually at l<strong>east</strong> (5–)7 mm long in flower, <strong>to</strong> 25 mm long<br />

with age; leaf sheaths and rhizomes usually purplish red; distal leaf sheaths with<br />

apex firm and opaque, nei<strong>the</strong>r inflated nor wrinkled, usually not disintegrating;<br />

<strong>plants</strong> <strong>to</strong> 115 cm tall.<br />

28. Culms round, internally hollow, completely and regularly septate with usually<br />

externally visible septa ca. 2–5 mm apart; scales <strong>of</strong> spikelet 100–500+ per<br />

spikelet, 1.5–2.7 mm long, 1–1.5 mm wide; achenes 0.7–1.1 mm long, <strong>the</strong> surface<br />

with a distinct finely reticulate pattern; tubercle 0.2–0.35 mm long; distal<br />

leaf sheath apex with a distinct <strong>to</strong>oth __________________________________ E. montana<br />

28. Culms round <strong>to</strong> conspicuously flattened, spongy internally; scales <strong>of</strong> spikelet<br />

15–100 per spikelet, 2–5.5 mm long, 1.5–2.5 mm wide; achenes (0.9–)1.1–1.9<br />

mm long, <strong>the</strong> surface � smooth in appearance; tubercle 0.3–0.7 mm long;<br />

distal leaf sheath apex with OR without a distinct <strong>to</strong>oth.<br />

29. Lowermost scale <strong>of</strong> spikelets clasping 2/3(–3/4) <strong>of</strong> culm; all spikelets with<br />

scale above <strong>the</strong> lowermost empty; culms round or slightly compressed<br />

_____________________________________________________________ E. palustris<br />

29. Lowermost scale <strong>of</strong> spikelets clasping 3/4 <strong>to</strong> all <strong>of</strong> culm; scale above <strong>the</strong><br />

lowermost with flower or empty; culms round <strong>to</strong> markedly compressed.<br />

30. Scale above <strong>the</strong> lowermost scale <strong>of</strong> some spikelets empty; lowermost<br />

scale <strong>of</strong> some or all spikelets usually clasping less than all <strong>of</strong> culm;<br />

culms round <strong>to</strong> markedly compressed; an<strong>the</strong>rs 1.3–2.7 mm long<br />

____________________________________________________ E. macrostachya<br />

30. Scale above <strong>the</strong> lowermost scale <strong>of</strong> all spikelets with flower; lowermost<br />

scale <strong>of</strong> all spikelets clasping all <strong>of</strong> culm (amplexicaulous); culms<br />

round; an<strong>the</strong>rs 1–1.8 mm long ______________________________ E. erythropoda<br />

22. Achenes trigonous (= 3-angled), sometimes bluntly so, <strong>to</strong> nearly rounded (definitely not<br />

biconvex) in cross section; styles 3-branched; stigmas 3.<br />

31. Tubercle huge, ca. as long and wide as achene; achenes coarsely honey-combed<br />

reticulate ____________________________________________________________ E. tuberculosa<br />

31. Tubercle much smaller than achene; achenes smooth <strong>to</strong> finely reticulate <strong>to</strong> coarsely<br />

honey-combed reticulate.<br />

32. Surface <strong>of</strong> achenes with a number <strong>of</strong> distinct ribs or ridges running lengthwise,<br />

with fine horizontal lines between <strong>the</strong> ribs ____________________________________ E. wolfii<br />

32. Surface <strong>of</strong> achenes smooth, without longitudinal ribs or ridges.<br />

33. Achenes widest at apex, truncate, black or nearly so, with <strong>the</strong> tubercle flattish<br />

across <strong>the</strong> entire apex (sometimes slightly wider than <strong>the</strong> achene) and with a<br />

small central point; apex <strong>of</strong> distal leaf sheath truncate, with an abrupt awl-shaped<br />

<strong>to</strong>oth; spikelets sometimes proliferating ____________________________ E. melanocarpa<br />

33. Achenes not widest at apex, not truncate, variously colored, <strong>the</strong> tubercle narrower,<br />

not stretching across <strong>the</strong> entire apex; apex <strong>of</strong> distal leaf sheath various;<br />

spikelets not proliferating except in E. rostellata and E. vivipara.<br />

34. Achenes constricted at summit <strong>to</strong> form a distinct narrow neck 0.2–0.3<br />

mm long; species rare in East TX, known only from Hardin Co. ___________ E. elongata<br />

34. Achenes variously shaped, <strong>of</strong>ten somewhat narrowed apically, but without<br />

a distinct narrow neck; including species widespread and common<br />

in East TX.

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